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Bioinformatics and Computational Biology (PhD)

Program description.

Researchers in the field of bioinformatics and computational biology collect, store, analyze, and present complex biological data using high-performance computing. Through this work, critical contributions are made to disease detection, drug design, forensics, agriculture, and environmental sciences. This research-oriented program trains a new generation of computational biologists for careers in private industry, academia, and government agencies. The program provides students with interdisciplinary academic curriculum that includes fundamental bioscience courses as well as advanced courses in bioinformatics.

In general, course requirements are completed within the first two years of study. Completion of coursework, the comprehensive exam, and a successful dissertation proposal results in advancement to candidacy status. In the next phase, students conduct research guided by a faculty member. The program is concluded with the defense of PhD dissertation.

The program is structured to be accessible for full- and part-time students. The courses are offered in a hybrid or distance learning format without requiring students to travel to campus. Because the research work is computational, students may choose to earn their doctoral degree entirely remotely or in a traditional face-to-face format.

What it costs

The total cost for this program is  $793.25  per credit hour for VA state residents or  $1,681.25 per credit hour for Non-VA residents. New Student and other mandatory university and course fees may apply.

Application Deadline

For application information and deadlines, see the Admissions site .

Program Summary

100 Percent Online

Program: Bioinformatics and Computational Biology

Degree: PhD

College/School: College of Science

Credits: 72 total

 Who should apply?

The program is inherently interdisciplinary and consequently the students coming to the program are likely to have a background in biology, computer science, or chemistry. Students select one of the areas in bioinformatics and computational biology studied by a faculty member, who will serve as their advisor.  Thus, the students interested in studying biological problems utilizing computational methods are encouraged to apply.

Why choose George Mason?

• Life science is a rapidly developing field demanding highly trained computational biologists. Our program addresses the growing demand by preparing a new generation of bioinformaticians to be employed in industry, academia, or government.
• By joining the program, students will participate in advanced, cutting-edge research.
• Students benefit from cooperation with neighboring world-class research institutions, including the NIH, USDA, FDA, FBI, and many other universities, research institutes, museums, government, and military laboratories in the Greater Washington, DC area.
• Courses in this program are taught by acclaimed professors and experts in the field, including Saleet Jafri, Dmitri Klimov, Donald Seto, Jeffrey Solka, Aman Ullah, and Iosif Vaisman as well as faculty from other departments.
• The program offers excellent opportunity for professionals with MS degree to complete PhD program on a part-time basis.
• The entire doctoral degree can be earned online without the need to travel to campus.

Program requirements are subject to change; please refer to the Mason Course Catalog for the most up-to-date requirements.

Requirements: 72 credits total

• 6 credits fundamental biosciences courses
• 13 credits core courses
• 3 credits colloquium course (1 credit each instance)
• 3 credits lab rotation (1 credit each instance)
• 12-24 credits research course
• 23-35 credits elective courses

All the courses needed to complete the entire program are offered online. Most online courses are  delivered synchronously . Classes are scheduled 4:30 p.m. (eastern standard time) or later to accommodate students employed full-time. Offerings vary by semester.

Fundamental Biosciences Courses

• BINF 701 Systems Biology | 3 credits
• BINF 702 Biological Data Analysis | 3 credits

Core Courses

• BINF 690 Numerical Methods for Bioinformatics | 3 credits
• BINF 705 Research Ethics | 1 credit
• BINF 730 Biological Sequence and Genome Analysis | 3 credits
• BINF 731 Protein Structure Analysis | 3 credits
• BINF 740 Introduction to Biophysics | 3 credits

Colloquium (3 instances)

• BINF 704 Colloquium in Bioinformatics | 1 credit

Bioinformatics Lab Rotation (3 instances)

• BINF 703 Bioinformatics Lab Rotation | 1 credit

Dissertation Research

• BINF 998 Doctoral Dissertation Proposal | 1-12 credits
• BINF 999 Doctoral Dissertation | 1-12 credits (must earn minimum 3 credits)

Electives may be graduate level coursework selected from bioinformatics, biology, biotechnology, statistics, computer science, and information systems courses. Review class schedule for latest course offerings. Contact your faculty advisor for approval of elective course selection.

Tuition (2021-2022)

Please Note : In addition to the tuition and fees described above, various course and lab fees may be assessed depending on the course(s) in which the student is enrolled. Please refer to the Students Accounts Office  website for more information on tuition and fees.

For information on loans and scholarships, visit the  Office of Student Financial Aid . For information regarding grants, tuition waivers and other merit aid, please inquire with your graduate department.

Career Descriptions

Because of explosive growth in life sciences, biotechnology, and drug design, there is a strong demand for bioinformatics scientists employing computational methods to advance the scientific understanding of living systems. Broadly speaking, the bioinformatics employment falls into several categories.

Researchers: Academia, government, and commercial sector are hiring employees trained in bioinformatics to support their research. People in these positions generally work in a specific area of research.

Instruction: There is a robust demand for teaching bioinformatics and computational biology. Some PhD level bioinformaticians will pursue an academic career developing their own research agenda and teach at universities. To this end, they typically receive an additional post-doctoral training in one of research labs for few years. Many institutions are also seeking dedicated instructors to teach bioinformatics.

Support of core facilities: Universities or government institutes tend to establish central resources for their labs referred to as core facilities. The personnel supporting such core facilities often require research experience and PhD level degrees.

Software developers: Many companies within the Greater Washington area and nationally are interested in developing and deploying computational algorithms and tools. Such jobs often require research experience and advanced skills only provided at PhD level.

Students are strongly encouraged to check online postings for job opportunities in bioinformatics and computational biology. As a Mason student, you may also contact  Mason Career Services  for more career information and counseling.

All applicants should review School of Systems Biology Application Information . For international applicants, see guidelines for  International Graduate Requirements.

Prerequisites

• Hold a bachelor’s degree in biology, computer science, or related field
• Achieve a minimum GPA of 3.25 in the last earned degree
• Have taken courses in molecular biology, biochemistry, genetics, calculus, computer programming and data structures, and probability and statistics. Students with deficiencies in one or more of these areas may be admitted, but required to take additional courses, some of which may not be applicable to the degree’s course total.
• Have a working knowledge of a computer programming language

Application Materials

• Graduate application
• Application fee
• In-state tuition form (if applicable)
• Two official transcripts from all colleges and universities attended
• Goals statement
• Three letters of recommendation from faculty members or individuals who have personal knowledge of your academic or professional capabilities

If you have any questions during the application process, please contact COS Graduate Admissions at [email protected] or (703) 993-3430.

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Ph.D. in Applied Biology

The Ph.D. Program in Applied Biology at UMass Lowell trains outstanding students to contribute to the wide array of life science research needs.

At a Glance

• Flexibility Part Time | Full Time
• Delivery Options On Campus

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UMass Lowell's Ph.D. in Applied Biology program integrates foundational areas of biology (e.g., Biochemistry, Cell & Molecular Biology, Genetics, Developmental Biology, Evolutionary Biology) with focused interdisciplinary biological research and preparation for career advancement.

The program combines fundamental and rigorous training in the pursuit of new knowledge regarding the mechanisms of biological processes coupled with the skills needed to apply this knowledge in the workforce. In addition to traditional academic careers in research and education at colleges and universities and academic research hospitals and institutes, our graduates are also prepared for non-academic careers.

To provide students an area of focus within the broad field of biology, the Applied Biology program includes four options. 

• Biomedical Science 
• Developmental & Evolutionary Biology 
• Quantitative Biology & Biophysics 
• Cellular & Molecular Biology 

Students select their option anytime during their first two years in the program. Each option contains a list of approved advanced option graduate courses. Students are required to complete at least four advanced courses in their option and two courses outside of their option.

For more information on the curriculum including course listings and degree pathways, visit the Graduate Catalog .

Admission Requirements

Specific aspects of admission to the Applied Biology Program are described below. These policies are in addition to all the standard requirements for graduate admission described by UMass Lowell Graduate Admissions office.

Faculty Sponsorship

Admission to the Applied Biology Ph.D. program is contingent on securing a faculty sponsor. It is the responsibility of the applicant to identify potential advisors, communicate with them, and secure a letter of sponsorship from at least one. Letters of sponsorship should be uploaded with your application, and are separate from your three letters of recommendation. Applications that lack a letter of sponsorship are not considered complete, and will not move on to the next round of consideration. Many faculty have their own process for selecting among prospective Ph.D. students, so it is best to contact them early in your application process. Students will have the opportunity to select a different permanent advisor during year 1, but it is anticipated that one of the predetermined potential advisors will serve as the permanent dissertation advisor.

The Graduate Record Exam (GRE) is not required for admission, but may be requested by a faculty sponsor during the evaluation process. The TOEFL (Test of English as a Foreign Language), IELTS, or Duolingo English Test are required from international applicants that come from countries where English is not the primary (official) or second predominant language. A minimum score of 90 is required on the TOEFL.

Personal Statement

A personal statement must be submitted that describes the student’s educational background, research experience, and career goals. The student must identify at least one potential faculty advisor in the statement. Particular attention will be given to students who have research experience and display a commitment to pursuing excellence in science.

Letters of Recommendation

Three letters of recommendation from professional sources must be submitted at the time of application. Sources may include academic professionals or supervisors/colleagues from a current or previous employer. 

Application Deadline

February 15, applications received after 2/15 will be considered depending on RA/TA availability.

See the UMass Lowell online Graduate Catalog for additional requirements of the program.

Visit the Graduate Admissions website for more information about the application process.

Tuition & Financing

To facilitate research and academic achievement, the Department of Biological Sciences offers at least two years of teaching assistant support for Applied Biology students. Support includes tuition, fees and a stipend for living expenses. 

In subsequent years support funding can be supplied by the department, training programs and faculty research grants. Applied Biology is a multi-department program. Additional grants, scholarships or research support opportunities may be available through individual departments/programs. Applicants are strongly encouraged to have established contact with potential research mentors before applying.

Breakdown of tuition and fees

Cost Per Credit: $842.21*

OUT-OF-STATE/INTERNATIONAL STUDENTS

Cost Per Credit: $1,496.66*

* These estimated graduate charges are for the 2019-2020 academic year. These prices are subject to change by vote of the UMass Board of Trustees.

• Reduced tuition programs : includes scholarships, financial aid and paid internships for qualifying students
• Free Application for Federal Student Aid (FAFSA) : U.S. residents are eligible to file to qualify for loans and other grant programs

For more information on tuition and financing, visit  Costs & Aid for Graduate Programs .

Why study Applied Biology at UMass Lowell?

Experiential Learning

Students have the opportunity to experience laboratory rotations and engage in journal clubs and research seminars. Students also work closely with a faculty advisor in the development of an individual research development plan that will be tailored to serve specific research and professional goals. Learn more about research topics

Cutting-edge Research

Assoc. Prof. Jessica Garb and a team of researchers including UMass Lowell students are examining the molecular composition, biomechanical properties and evolution of an exceptional type of silk produced by an arachnid species known as Darwin’s bark spider. Read more

Outstanding Faculty

Asst. Prof. Jennifer L. Fish is spearheading a project to understand the molecular mechanisms responsible in patients who have mutations in the gene called SATB2and how these mutations in SATB2 cause defects, specifically in the growth of the lower jaw. Read more

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Temple University

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Biology PhD

Gain the experience to make significant contributions to biological research with the Biology PhD program in the College of Science and Technology. The Biology PhD degree provides a rigorous, research-focused course of study. You’ll be prepared to pursue multiple career pathways related to the biological sciences, including postdoctoral and academic positions, and in related industries, such as biotechnology, bioengineering, healthcare and pharmaceutics. The Biology PhD program provides opportunities for you to receive funding for your degree with either a teaching or research assistantship, and a tuition waiver.

Biology PhD students are afforded the chance to choose which lab they want to join. During your first year, you can choose a specific research group immediately or you can rotate up to three research groups before deciding on a lab. Once you are admitted to candidacy, you’ll pursue specialized research toward the completion of your degree.

Faculty members perform cutting-edge research in different areas of specialization that include

• cellular, developmental and molecular biology;
• cellular and molecular neuroscience, which investigates the central nervous system and the mechanisms it governs;
• computational biology, which focuses on computational evolutionary genomics;
• ecology, an interdisciplinary field that investigates the complex relationships between living organisms and their environments;
• evolutionary medicine, an interdisciplinary field that examines contemporary human disease within the larger context of evolutionary history;
• genetic epidemiology, an interdisciplinary field that examines the impact of genetic and environmental factors on modern human disease; and
• population genetics, a subfield of genetics and evolutionary biology that examines genetic change over time within and between populations.

Learn more about the Department of Biology’s research areas .

Coursework is interdisciplinary and students are encouraged to participate in research across other fields, including biochemistry, chemistry, computer science, ecology, environmental science, engineering, mathematics and physics. Doctoral candidates perform innovative research in many of the College of Science and Technology’s state-of-the-art centers and facilities, including the

• Center for Biodiversity , focused on the conservation, education and research of species and their impact on the environment;
• Center for Computational Genetics and Genomics , whose mission is to bring together scientists who work on genetic and evolutionary theory and who employ computational approaches; 
• Institute for Genomics and Evolutionary Medicine , focused on molecular evolutionary knowledge and tools to advance research and for product development in biomedicine and biodiversity
• Sbarro Health Research Organization , where genetic research investigates treatments and cures to chronic illnesses such as cancer, cardiovascular disease and diabetes.

Learn more about the Department of Biology’s research centers and facilities .

Program Requirements

Doctoral candidates have seven years to complete the required courses and examination, as well as a dissertation. All candidates must fulfill the following requirements for successful completion of the Biology PhD program. 

• an original research proposal , a 15- to 20-page proposal that introduces a research problem and the student’s methodological plan for investigating the problem, and
• the oral examination , which is administered after a student’s research proposal has been approved and tests a student’s understanding of the background and substance of the research proposal. (If a student’s research proposal is not approved, the student is considered to have not passed the oral examination.)
• The doctoral dissertation is an original, empirical study that demonstrates a student's knowledge of research methods and mastery of the primary area of research. All dissertation research is conducted under the guidance of the Doctoral Advisory Committee.
• The final dissertation defense is the culminating event for Biology PhD candidates. Students must successfully pass the dissertation defense for completion of the Biology PhD degree.  

All Biology PhD candidates must have experience teaching at Temple for a minimum of two semesters. This requirement can be satisfied through teaching assistantships in the Department of Biology.

Learn more about the Biology PhD examination and dissertation requirements . 

Classes & Curriculum

Following coursework in the first one to two years, you'll advance to candidacy and engage in research opportunities across the many labs and disciplines represented by faculty members. Doctoral students will undertake a preliminary examination, as well as an original dissertation, to demonstrate their knowledge of research methods and mastery of their primary research area.

The Biology PhD curriculum requires 36 credit hours, including 15 hours of graduate seminar courses. Some of these courses include

• Analytical Biotechnology , 
• Cellular and Molecular Neuroscience , 
• Contemporary Biology , 
• General Biochemistry, and 
• Evolutionary Genetics Genomics . 

Learn more about required courses for the Biology PhD .

Related Graduate Degrees

• Bioinformatics PhD
• Biotechnology PSM
• Scientific Writing PSM
• Bioengineering PhD *

*College of Engineering

Related Graduate Certificates

• Bioinformatics
• Bioinnovation
• Biotechnology
• Scientific Writing

Tuition & Fees

In keeping with Temple’s commitment to access and affordability, this Doctor of Philosophy offers a competitive level of tuition with multiple opportunities for financial support.

Tuition rates are set annually by the university and are affected by multiple factors, including program degree level (undergraduate or graduate), course load (full- or part-time), in-state or out-of-state residency, and more. These tuition costs apply to the 2023–2024 academic year.

Pennsylvania resident : $1,353.00 per credit Out-of-state : $1,765.00 per credit

You can view the full Cost of Attendance breakdown on the Student Financial Services website .

Faculty advisors can help you navigate your program and discuss research opportunities. There are two graduate advisors for the Department of Biology.

Tonia Hsieh  is an associate professor and the graduate chair in the Department of Biology. Phone : 215-204-0617 Email : [email protected]

Sandhya Verma is the graduate administrator in the Department of Biology. Phone : 215-204-8854 Email : [email protected]

If you can find the time to participate while meeting the demands of your degree program, student clubs offer valuable opportunities to meet fellow graduate students, researchers and professionals from science- and technology-related industries.

Clubs and organizations for students pursuing a Biology degree include

• the Alliance for Minority Participation in STEM , which works to increase opportunities for students and help them complete their undergraduate and graduate degrees in STEM.
• the Biology Graduate Student Society , an organization that develops professional and leadership skills in doctoral students as well as encourages them to participate in public outreach.
• Leadership, Education and Development in Science , an organization that empowers Philadelphia middle school students to form an interest in STEM through science experiments and mentors.

Explore the full list of College of Science and Technology student clubs and organizations .

If the demands of your degree program allow it, consider spending a summer or semester overseas building international connections with researchers from other countries. Temple’s study abroad program offers College of Science and Technology students opportunities to spend a summer or semester studying in China, Germany, Italy, South Korea or Taiwan. Learn more about your study abroad options .

As a division within Temple University, the College of Science and Technology is accredited by the Middle States Commission on Higher Education .

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Doctor of Philosophy (PhD)

Offered By: Department of Biochemistry and Molecular Biology

Onsite | Full-Time | 5 – 6 years

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About the PhD in Biochemistry and Molecular Biology Program

In the Biochemistry and Molecular Biology PhD program, faculty, and students work together to increase knowledge of the biochemical and molecular bases of normal and abnormal cellular processes. Our program trains students to be successful independent scientists and gives them the knowledge, research training, and leadership skills to continue to provide new insights into the biomedical issues that have a profound impact on public health.

Students engage in a rigorous course curriculum and a range of structured and informal activities outside the classroom and lab to build their skills. They will pursue their thesis research in the lab of one of our over forty training faculty across the Johns Hopkins Bloomberg School of Public Health and the Johns Hopkins School of Medicine.

Visit our dedicated PhD program website to learn more about the diverse research training opportunities of the program.

PhD in Biochemistry and Molecular Biology Program Highlights

Our position within the School of Public Health provides a unique setting in which students learn how biochemistry, molecular biology, physical chemistry, cell biology, and genetics can be used to solve significant problems in public health and medicine. Our program offers:

• Training faculty from across the School of Public Health and the School of Medicine
• A strong grounding in the science of biomedical and public health research through a core curriculum that includes courses taught by leading experts from the Schools of Public Health and Medicine
• Training outside the lab and classroom in key skills such as communications and leadership
• Opportunities to build strong communications skills through a range of speaking venues including journal club, research colloquium, department retreats, and national meetings
• Access to the Johns Hopkins School of Medicine Professional Development and Career Office , offering excellent career services and professional development,  including the BMB-required OPTIONS program, a guided process of career exploration for paths from medicine to biotech to academia and beyond
• Opportunities to participate in community service and outreach, with a focus on our East Baltimore neighborhoods, through the Johns Hopkins University community engagement and service-learning center, SOURCE

Training faculty across the School of Public Health and the School of Medicine

Schools that students can take courses in: Public Health, Arts & Sciences, Medicine, and Engineering

Two-month rotations in the first year prior to selecting thesis lab

Average number of incoming students in the BMB PhD degree program each year

What Can You Do With a PhD In Biochemistry And Molecular Biology?

The Biochemistry and Molecular Biology PhD program prepares students for a range of biomedical and health sciences careers, including in academia, industry, policy, and beyond. Visit the Graduate Employment Outcomes Dashboard to learn about Bloomberg School graduates' employment status, sector, and salaries.

Sample Careers

• Research Scientist
• Science Policy Adviser
• Biotech Executive
• Senior Scientist
• Patent Lawyer
• Science Policy Analyst/Advocate
• Science Writer/Journalist
• Biological Sciences Teacher

Topic Areas

The BMB PhD program faculty conduct research to gain new insights into the cellular and molecular mechanisms underlying normal and abnormal cellular processes, and their relevance as targets for improving health and treating disease. Our training program places particular emphasis on mechanistic approaches to research problems.

Common topic areas within our faculty's diverse research interests include:

• Biophysics and Structural Biology
• Cancer Biology
• Chemical Biology and Proteomics
• Cell Biology
• Cellular Stress and Cell Signaling
• Genetics, Genomics, and Gene Regulation
• Immunology and Infectious Diseases
• Translational Research

Curriculum for the PhD in Biochemistry and Molecular Biology

The BMB PhD offers students a rigorous course curriculum, including a set of common core classes from the Schools of Public Health and Medicine. A rich array of seminar programs and journal clubs are also available to all students.

Browse an overview of the requirements for this PhD program in the JHU  Academic Catalogue  and explore all course offerings in the Bloomberg School  Course Directory .

Courses in core curriculum

Minimum elective credits

Seminars on current research presented by experts from across Johns Hopkins and other biomedical research institutions

Courses available across Johns Hopkins Schools of Public Health, Medicine, and Arts and Sciences

Admissions Requirements

For the general admissions requirements see our How to Apply page. The specific program also requires:

Prior Work Experience

Laboratory research experience (from academia, industry, etc.) is required

Prior Coursework

Strong background in the sciences, particularly in chemistry, biochemistry, or biology

Standardized Test Scores

Standardized test scores (GRE) are optional for this program. The admissions committee will make no assumptions if a standardized test score is omitted from an application, but will require evidence of quantitative/analytical ability through other application components such as academic transcripts and/or supplemental questions.  Applications will be reviewed holistically based on all application components.

Program Faculty Spotlight

Ashani T. Weeraratna

Ashi Weeraratna, PhD, studies how cancer cells move to distant sites and how changes in the normal cells around a tumor contribute to their movement, especially as we age.

Michael J. Matunis

Michael Matunis, PhD, studies how protein modification by SUMO—the small ubiquitin-related modifier—drives changes in key cellular pathways from stress response to DNA repair.

Jennifer M. Kavran

Jennifer Kavran, PhD, MS, MPhil, is a biophysicist who investigates how cells communicate with each other and their environment.

Danfeng Cai

Danfeng Cai, PhD, combines advanced microscopy, genomics, and proteomics to tease out the functions of protein condensates in cells, with a focus on cancer.

Vivien Thomas PhD Scholars

The  Vivien Thomas Scholars Initiative (VTSI)  is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically Black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. To be considered for the VTSI, you will need to submit a SOPHAS application, VTSI supplementary materials, and all supporting documents (letters, transcripts, and test scores) by December 1, 2024. VTSI applicants are eligible for an application fee waiver , but the fee waiver must be requested by November 15, 2024 and prior to submission of the SOPHAS application.

Per the Collective Bargaining Agreement (CBA) with the JHU PhD Union, the minimum guaranteed 2025-2026 academic year stipend is $50,000 for all PhD students with a 4% increase the following year. Tuition, fees, and medical benefits are provided, including health insurance premiums for PhD student’s children and spouses of international students, depending on visa type. The minimum stipend and tuition coverage is guaranteed for at least the first four years of a BSPH PhD program; specific amounts and the number of years supported, as well as work expectations related to that stipend will vary across departments and funding source. Please refer to the  CBA to review specific benefits, compensation, and other terms.

In the BMB PhD program, all full-time PhD students who remain in good academic standing will receive the above support through the entire duration of the program.

Need-Based Relocation Grants Students who  are admitted to PhD programs at JHU starting in Fall 2023 or beyond can apply to receive a need-based grant to offset the costs of relocating to be able to attend JHU.   These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their PhD program. This is not a merit-based grant. Applications will be evaluated solely based on financial need.  View more information about the need-based relocation grants for PhD students .

Questions about the program? We're happy to help.

Mike Matunis, PhD PhD Program Director

Roza Selimyan , PhD BMB Executive Director for Academic Affairs and Education Programs

Erika Vaitekunas Administrative Specialist

[email protected]

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Ph.d. biology.

Our PhD program accepts students possessing Bachelors or Masters degrees who are planning a career in science. The purpose of our PhD program is to prepare highly qualified scientists who have excellent up-to-date training in the fields of biology or bioinformatics, who are actively involved in scientific research, are capable of making significant contributions to their scientific field, possess all the necessary skills for effective oral and written communication with colleagues, and can successfully compete in the job market for postdoctoral and research scientist level positions in academia and science-related industries.

We currently have approximately 100 PhD students in the Biology PhD program of the School of Biological Sciences. Of recent PhD graduates, 70% are currently employed as postdoctoral researchers in academia, 9% as government scientists , 9% as industry scientists , 4% as instructors , and 9% are enrolled in further professional training . The average number of journal publications is 3.9 per student and the average number of presentations at conferences around the world is 4.3 per student .

Areas of concentration:

• Ecology, Evolution, and Behavior   including population and evolutionary ecology; community ecology; aquatic chemical ecology; ecological genomics; sensory ecology; evolution of development, behavior, and sociality; biological oceanography; environmental microbiology; theoretical ecology.
• Molecular and Cell Biology   including eukaryotic and prokaryotic cell biology; molecular physiology; molecular biophysics and structural biology; animal, plant, and microbial molecular genetics; human genomics; molecular evolution.

Quick Facts

• Stipend and tuition waiver offered.
• Fellowships available.
• Duration of program depends on research progress. Ph.D. candidates typically defend their thesis at the end of the 5th or during the 6th year.
• 18 credit hours of coursework.

General Inquiries

• Chung Kim Academic Program Coordinator Email  | 404.385.4240 | EBB 2009

Ph.D. Program Overview:

• Stipend: $35,500 per year
• Year 1 Focus:  Coursework / Lab Rotations / Teach
• Year 2 Focus:  Coursework / Thesis Research / Qualifying Exams
• Year 3 Focus:  Thesis Research
• Year 4 Focus:  Thesis Research
• Year 5 Focus:  Thesis Research, Writing and Defense

The PhD in Biology is a research-based degree involving deep immersion in a topic within biology with mentoring from an advisor and thesis committee with expertise in the field of study. Coursework is typically completed within the first 2 years (18 credit hours). Students who have previously earned an M.S. degree or taken graduate courses elsewhere can negotiate up to 9 transfer credit hours upon entering our program. Selection of a thesis advisor is made in discussion with our faculty and/or via lab rotations during the first year, and the composition of the thesis committee is established by the student by the end of the first year.

PhD students in the School of Biological Sciences are each supported by a stipend and do not pay tuition, only modest   fees   each semester. Stipend support comes from teaching or research assistantships which complement research training towards the PhD. Whether PhD stipends are earned from research or teaching assistantships after the first year is decided in consultation with a student's thesis advisor and the graduate committee. Our base PhD stipend is $35,500. Several fellowships are available which can supplement the PhD stipend.

Further Funding

Please take some time to review our   funding opportunities and fellowships .

Choosing between the Masters and Ph.D. Program

Our graduate program is primarily focused on the PhD which prepares students for careers in scientific research and employment in academia, industry, or government. We encourage potential applicants to the PhD program to directly contact individual   faculty   members whose research may appeal to them to discuss research interests and future opportunities. Students who are not yet sure of their interest in scientific research or are interested in other kinds of professional development should consider the M.S. degree in   Biology   or  Bioinformatics . Admission decisions are made by our graduate committee (composed of Biology faculty) in consultation with all of the faculty in the School of Biological Sciences.

Georgia Tech provides application fee waivers to advance diversity, recognize outstanding undergraduate achievement, and engage prospective students in recruitment events where Georgia Tech is affiliated. Such fee waivers are currently available only to domestic applicants.

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Biology related research centers.

• Aquatic Chemical Ecology Center
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• Center for NanoMAD
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• Biology (Ph.D.)

Prepare for a career in research or academia through a combination of extensive research and coursework to earn a Ph.D. in biology at Illinois Tech.

• Academic Programs

Illinois Tech's Ph.D. program in biology is tailored to fit your background and goals. Programs of study may be designed in any of the three areas of concentration: biochemistry, cell and molecular biology, or microbiology. The department has state-of-the-art computer and laboratory equipment in which to conduct research in these areas.

In addition, the department constructs and operates facilities for X-ray scattering and imaging at the Advanced Photon Source at Argonne National Laboratory. The department offers graduate programs leading to Ph.D. degrees in biology, concentrating educational and research activities in the areas of biochemistry, bioinformatics, cell and molecular biology, and microbiology. Graduate education in biology is available on either a full- or a part-time basis.

Program Overview

The Ph.D. program is tailored to fit your background and goals. The department has state-of-the-art computer and laboratory equipment in which to conduct research, including X-ray scattering and imaging at the Advanced Photon Source at Argonne National Laboratory.

Career Opportunities

A career with a Ph.D. in biology will depend on the concentration of study.

• Microbiologist
• Environmental scientist
• Medical scientist
• Natural sciences manager
• Postsecondary biology teacher

View details

Admission Requirements

Completion of an M.S. degree is not normally required for admission to the full-time Ph.D. degree program, but may be required of part-time students.

You must pass the Ph.D. qualifying examination in your respective area of specialization: biochemistry, cell and molecular biology, or microbiology.

• GRE: 310 (quantitative and verbal), 3.0 (analytical writing)
• GPA: 3.0/4.0

Informing Agricultural Practices Through Computational Genomics

Biology Ph.D. Candidate Dakota Bunn investigates the habits of the western bean cutworm, a pest to corn and dry beans, using computational genomics.

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In the field of bioinformatics and computational biology, specialists collect, store, analyze, model, and present complex biological data to understand how individual components of biological systems and the interactions between them inform function at all levels of biological hierarchy. Through the combination of biological analysis and high-performance computing, critical contributions are made to biomedical, agricultural, and environmental sciences. The main objective of this doctorate is to educate the next generation of computational biologists for careers in academia, industry, and government. The program provides students with interdisciplinary academic training that includes fundamental bioscience courses as well as advanced courses in bioinformatics, which cover the most recent developments in the field. The curriculum is designed to be completed in approximately two years. The completion of coursework, the comprehensive exam, and a successful dissertation proposal result in advancement to candidacy status. In the final phase of the program, students focus on research that culminates in a dissertation.

The program is structured to be accessible for full and part-time students. Because the program has a fully online option, students may choose to earn the doctoral degree entirely remotely or in a traditional face-to-face format. All courses are offered in hybrid, distance learning, or asynchronous formats without requiring students to travel to campus. Visit Mason Online for details.

University-wide admissions policies can be found in the  Graduate Admissions Policies  section of this catalog. International students and students having earned international degrees should also refer to  Admission of International Students  for additional requirements.

Eligibility

Applicants should have a bachelor's degree in biology, computer science, or a related field from an institution of higher education accredited by a Mason-recognized U.S. institutional accrediting agency or international equivalent with a minimum GPA of 3.25 in the last earned degree. Applicants are expected to have completed coursework in molecular biology, genetics, calculus, chemistry, computer programming, and probability and statistics. Students with deficiencies in one or more of these areas may be admitted and required to take additional courses, some of which may not be applicable to the degree's credit total.

Application Requirements

To apply for this program, prospective students should submit the  George Mason University Admissions Application  and its required supplemental documentation, a goals statement, and three letters of recommendation.

The GRE is not required for admission into this program.

For policies governing all graduate programs, see  AP.6 Graduate Policies .

Transferring Previous Graduate Credit into this Program

Previously earned and relevant graduate credits may be eligible for transfer into this program; details can be found in the  Credit by Exam or Transfer  section of this catalog.

Banner Code: SC-PHD-BCB

Degree Requirements

Total credits: 72

Students should refer to the Admissions & Policies tab for specific policies related to this program.

Doctoral Coursework

Doctoral Committee and Advancement to Candidacy

By the end of the semester when coursework is completed, the student must form a doctoral committee made up of a minimum of three graduate faculty members and take a written comprehensive exam. The exam includes written and oral components. Upon passing the comprehensive exam and submitting an acceptable dissertation proposal, the student is advanced to doctoral candidacy to begin the dissertation writing phase.

Dissertation Research

A minimum of 12 and maximum of 24 combined credits from BINF 998 Doctoral Dissertation Proposal and BINF 999 Doctoral Dissertation may be applied toward satisfying doctoral degree requirements. Students must take at least 3 credits of BINF 999 Doctoral Dissertation .

Doctoral Dissertation

After advancing to doctoral candidacy, students work on their doctoral dissertation while enrolled in BINF 999 Doctoral Dissertation . The dissertation should represent a significant contribution that is suitable for publication in a refereed scientific journal. The dissertation must be defended in a public forum before the dissertation committee and other interested faculty.

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Best Part-Time PhD and Doctoral Programs (Online & Campus)

Learn how you can earn your PhD or professional doctorate part-time, plus find top colleges offering these flexible programs online and on-campus.

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Are you eager to expand your expertise and become a leader in your field? Perhaps you have already obtained a bachelor’s or master’s degree and are looking to advance your career further. Pursuing a PhD or professional doctorate degree is the next logical step. However, you may be hesitant about taking on an intense doctoral program while also working full-time.

Fortunately, quality universities offer part-time programs (including convenient online options) that can make earning your doctorate much more manageable. This guide provides insight into the best part-time PhD and doctoral programs, explores common fields of study that offer part-time programs, and answers frequently asked questions about these programs. Keep reading to determine if a part-time doctoral program is right for you and find the one that can take your career to the next level.

Part-Time PhD and Doctoral Programs: Best in 2024

Quality, affordability, and accreditation are important when searching for the best part-time PhD and doctoral programs. The best choice for you, though, often boils down to how well a program fits with your full-time work schedule and everyday life. We’ve rounded up the top part-time PhD and doctoral programs that are flexible for maintaining work, life, and education balance while also providing a high-quality education.

Johns Hopkins University’s Part-Time Online DrPH

John Hopkins University provides public health professionals with a robust Doctor of Public Health (DrPH) . The program helps early- to mid-career professionals working in international or domestic public health roles become leaders in their fields. There are ten concentrations, and the program helps connect students with the right courses, opportunities, and research projects to reach their individual goals. 

This program offers part-time enrollment and can be completed in one year when attending full-time. To be eligible, applicants must hold an MPH from an accredited university and have at least three years of work experience in the field. Additionally, John Hopkins offers competitive scholarships to help offset the program’s costs.

George Mason University’s School of Policy and Government Part-Time PhD Programs

George Mason University offers three PhD programs – biodefense, public policy, and political science – to those working in the public policy sector. The Schar School PhD programs are designed for those who want a heavy research focus and the opportunity to conduct ground-breaking research in their fields. Alumni find success working in academia, for government organizations, and at nonprofits thanks to the school’s robust dissertation program, rigorous coursework, and networking opportunities. 

Each program is available part-time or full-time and offers opportunities to customize the curriculum to your career goals. Prospective students need previous educational and work experience in the public policy field.

Boston College’s Part-Time (Ed.D.) in Higher Education

Boston College’s Executive Ed.D. in Higher Education program exists for those looking to advance their careers in higher education. The program develops innovative leaders who are ready to tackle the big problems facing higher education today. A unique feature of this program is how the school designs cohorts so each student comes from a different background to increase the diversity of thought in the program. Courses are all offered online and at times that work around a typical nine-to-five career. 

Program requirements include summer residencies in Boston and Dublin. This three-year program also boasts a global focus with research and networking opportunities across the globe. Successful applicants need at least seven years of professional experience and a master’s degree.  

Wayne State University’s Part-Time PhD in Social Work

Wayne State University offers one of the best part-time doctoral programs in social work in the country for those who already have a master’s in social work. With options in anthropology, gerontology, or infant mental health emphasis, the first three years of coursework prepare students for the final phase of the program – a dissertation. Graduates of WSU’s doctoral programs often take on leadership roles in academia, clinical scholarship, and social welfare policy. While part-time enrollment is possible, students must complete the program within seven years.

Drexel University’s Part-Time Online DBA

The Executive Doctorate in Business Administration (DBA) program from Drexel University offers business professionals an online option to gain a high-quality doctoral education on their own schedules. The program prepares leaders for the business world and focuses on applying business theory to real-world scenarios. Students can also take PhD-level courses from other disciplines to integrate non-business ideas into their dissertations and projects. 

Drexel designed its program for those with over ten years of experience in business, and the admissions process is highly selective with each cohort consisting of only 12-18 students. While most PhD programs are completed in three years, this program can be completed in two and a half when enrolled full-time, though part-time enrollment is also an option. 

Most Popular Part-Time PhDs and Doctorates

Part-time PhD and professional doctorate programs are available in various fields of study. While professional doctorate programs emphasize applying research in practical settings and are commonly offered in part-time or online formats, PhD programs require original research contributions. Nevertheless, there are several popular part-time PhD and professional doctorate programs to choose from. The list below provides an overview of some of the most sought-after programs and can help you explore your options.

Criminal Justice

Since PhD and doctoral programs in criminal justice are for those already working in the criminal justice field, many are part-time and online . A PhD is more research-focused, like the Criminal Justice with a Concentration in Intelligence (Ph.D.) program at the University of North Georgia. A professional doctorate program, like the Doctoral Program in Criminal Justice at Texas State University, is focused on applying academic theories to professional settings. Many of these degrees are available online and on a part-time or full-time basis.

A doctorate or PhD program in nursing is a great option for nurses want to become nurse practitioners, teach at the college level, or conduct original research. Since candidates usually need master’s degrees and previous work experience, many online Doctor of Nursing Practice (DNP) programs are designed for working professionals. There are two types of advanced nursing programs commonly offered, the PhD in Nursing and a Doctor of Nursing Practice (DNP). The Doctor of Nursing Practice (DNP) at Baylor University and the online PhD program at George Washington University are examples of these types of advanced nursing programs.

A PhD in business or Doctorate of Business Administration (DBA) are some of the most common part-time programs designed to be completed while working full-time. Many online DBA programs and online PHD programs in business work especially well for busy professionals. The DBA at Washington University and the Katz Executive DBA from the University of Pittsburgh are examples of online, part-time options for advanced business degrees.

Education doctorates and PhD programs work well for teachers and educators wanting advanced leadership influence in the field. The programs are usually part-time and work around a typical academic schedule. Online education doctorate s focus on applying academic theories to educational settings, while online PhD in education programs are more focused on academia. The PhD program from Harvard University , available on campus and either part-time or full-time, and the part-time Doctor of Education – Online with Residency program from Northeastern University are examples of popular advanced education degree programs.

Communications

A doctorate or PhD program in communication is a great option for those already in the communication field who want to enhance their expertise. Since candidates often need a master’s degree and previous work experience in the field to apply, many online communications doctorate programs are designed for working professionals. The online PhD in Communication program at American University and Columbia’s PhD in Communications give examples of popular doctorate and PhD programs in education.

Public Health

A Doctor of Public Health is one of the most common part-time advanced programs and is designed to be completed while working full-time. Many online public health doctorates work especially well for working professionals, some with work requirements built into the curriculum. That means having a full-time role can actually help you progress more quickly through the program. Boston University and Penn State University both offer part-time PhD programs.

Key Questions About Part-Time PhD and Doctoral Programs

Are part-time phd and doctoral programs offered online.

While it is highly subject-dependent, many PhD and doctoral programs are offered online . However, it’s not only the subject matter that can determine if a program is offered online but the type of degree, too. Since a professional doctoral program is designed for working professionals, it is often offered online and around a typical work schedule. The Doctor of Public Health (DrPH) from John Hopkins University and the public policy PhD programs from George Mason University are examples of these types of flexible programs. However, PhDs are more academia focused and less likely to be offered online since they often have an in-person research component.

How long does it take to complete a part-time PhD?

The time it takes to complete a part-time PhD program depends on the curriculum, how many courses you take each semester, and the timeline for your dissertation. On average, expect a part-time PhD to take three to five years with some programs having a seven-year maximum you can spend on your coursework and dissertation.

Are part-time PhD students eligible for financial aid?

While PhD programs cost more than undergraduate and master’s programs, there’s also often more financial aid available to doctoral students. PhD students are often eligible for PhD-specific grants and scholarships as well as fellowship and tuition reimbursement opportunities. For more information on completing your PhD on a budget, check out our financial aid guide .

Am I still required to do a dissertation as a part-time PhD student?

Most part-time PhDs still have a dissertation requirement. Some professional doctorate programs have shorter research projects or capstones instead, which are less of a commitment than a dissertation. If you are looking for a program with a capstone instead of a dissertation, it’s best to talk directly to the programs you’re interested in about research requirements.

Do part-time doctoral programs have residency requirements?

This depends on your chosen field of study. Health-related fields and social work programs typically have residency requirements, but many programs outside of health and social work have them as well. For example, Boston College’s Executive EdD program has summer residency requirements for students. Residencies are often tailored around a working professional’s schedule, though, and work experience can sometimes be substituted in place of the requirement.

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5 Best Work-From-Home Jobs For PhDs

Written by Tavis Mendez, PhD

I love being a scientist.

But, getting up in the morning and heading into the lab has always been a hassle for me.

I am also quite introverted and prefer being by myself most of the time.

During my PhD, the days I enjoyed most were the ones when I could stay home and focus on writing articles and mining through data.

It was wonderful not having to stand at the lab bench all day.

But, as a scientist, wasn’t I supposed to love bench work?

Was I going to have to keep doing the bench work I hated if I wanted to stay in science?

It seemed like the only option for me after my PhD was to become a postdoc and do more bench work.

This idea caused me stress and anxiety .

I didn’t want to stay in academia, but I wasn’t sure what else to do.

I decided to go to a networking event to explore my career options outside of academia.

At the event, I met someone who was working in the sciences, at home, on her own schedule.

It sounded amazing.

I set up an informational interview with this new connection, and realized that there are many alternative careers available to PhDs, including working remotely.

It was clear to me now, I was going to leave academia and get an industry position where I could work from the comfort of my home.

I brushed up on my interviewing skills and crafted my industry resume .

Surprisingly, I managed to successfully network as a quiet introverted PhD and secure a job referral.

After a series of interviews, I was able to find a job perfectly suited to me.

My new job allowed me to work from home, doing what I love.

Why Work-From-Home Jobs Are A Great Option For PhDs

The Internet has changed, and continues to change, our society.

According to a study by CISCO, reported in The Huffington Post , there are currently 3 billion unique Internet users around the world and this number is expected to rise to 4.1 billion by 2020.

The Internet is a driving factor in economic growth, and as more and more people are using the Internet, its contribution to the economy will continue to increase.

But, not only is the Internet driving the economy, it’s changing the way people work.

According to the Bureau of Labor Statistics , 38% of people in leadership and managerial positions reported working remotely some or all of the time.

As a PhD, you are well-suited to leadership roles , making remote work a definite possibility for your future.

Working remotely is increasing in popularity, with many large companies offering telecommute opportunities.

A survey by Flexjobs found that 68% of respondents expect to work remotely in the future.

If working remotely appeals to you, there are plenty of options available now, and even more are likely to appear in the future.

But what PhD industry careers are the most well-suited for remote work?

Top Jobs For PhDs To Work From Home

A remote job gives you the freedom to choose where you work.

You could work from home, from a coffee shop, or from your latest travel destination.

Take the initiative to seek out these new opportunities and don’t stay stuck in academia , trapped in an office or lab.

If a flexible job is what you want, you already have the transferable skills to get your desired position.

While many jobs have the potential to be telecommute-friendly, a few stand out from the rest as the best options for PhDs.

Here are the top 5 industry positions that allow you to work from home…

1. Medical writing.

Medical writing is a profession that has gained a lot of steam in the past few years.

There are more biomedical studies now than ever before, and the need for PhDs with high quality communication skills to write about these studies is increasing.

As a medical writer, you will be expected to write about the topics requested by a client, and you will have to work to meet tight deadlines.

But, you won’t have to go to an office to do your work. All your writing can easily be done from home.

Another advantage of medical writing is that you will stay connected to new advancements in the medical field.

Each assignment will be different and you will need to learn new aspects of the medical industry.

Medical writers are truly life-long learners.

Whether you are crafting regulatory documents in clinical research, designing promotional literature, or writing content in a healthcare journal, a medical writer position will require your PhD research prowess and your creativity.

According to PayScale , the average salary for a medical writer is $71,000.

2. Scientific editing.

Scientific editing is another career that allows you to maintain a connection with cutting-edge research discoveries without having to spend all your time at the lab bench.

By editing scientific research papers, you will have access to new research in real-time.

In this position, you will organize peer reviews, as well as proofread and edit research papers.

Attention to detail, good time management, and high-level research and writing skills are essential for a scientific editor.

As a PhD, you have already developed these desired transferable skills .

There are many peer-reviewed journals, editing firms, and research institutes in need of PhDs to edit the growing number of research papers.

Editors are needed for all different areas of research.

You can always choose a subject that you are well acquainted with or something outside of your comfort zone.

As a PhD, you can learn anything, so do not feel limited to edit only in scientific areas where you have experience.

According to Glassdoor , the average salary for a scientific editor is $73,000.

3. Consulting.

In industry, there is a growing need for consultants who can provide businesses with expert advice.

As a PhD, you are an expert in your field and you have high-level learning skills that give you the ability to become an expert in other fields as well.

Consulting is a diverse career that includes scientific consulting, as well as legal, media, or corporate consulting.

The analytical and research skills you have as a PhD make you an ideal candidate for a consulting position.

While many consultants work for a consulting firm, it is also possible to start your own consulting business, which would give you more work flexibility.

As an independent consultant, you need excellent communication and networking skills to find and secure clients.

This is a great job for extroverted PhDs who enjoy networking and making new connections all the time.

An independent consultant also needs a strong entrepreneurial mindset, since you will need to successfully market and sell your services.

The average salary of a consultant varies depending on the field of expertise.

4. Translation.

Translation is an uncommon career choice, but it is where fluency in more than one language can be advantageous to the advancement of your career.

Every industry has its own unique technical language, especially the biomedical sciences, that can be difficult for non-scientists to understand.

If you understand scientific or technical lingo in more than one language, you are a great candidate for a career in translation.

Beyond the technical jargon, each culture has its own expressions and, as a translator, being able to express each sentence so it is understood in another language is crucial.

Most translators do not have a scientific background, but as a multilingual PhD you have an advantage over these non-scientists.

As diversity in the US increases and the global economy grows, the demand for translators is expected to increase.

Fluent understanding of more than one language is a great advantage when combined with the transferable skills gained during your PhD, making you a great job candidate.

According to US News , translators can earn up to $78,000 per year.

5. Senior clinical research associate.

The senior clinical research associate role is a great alternative industry PhD career , but it’s not normally associated with working remotely.

However, according to Forbes , 78% of senior CRAs report that they are able to complete their jobs remotely.

In the senior CRA role, you will be handling clinical trials and ensuring that every project follows Good Clinical Practice guidelines.

You will be expected to manage teams of people, create project outlines, and meet deadlines — all things you learned to do as a PhD student or postdoc.

The senior CRA is responsible for everything relating to the clinical study, including budget, training, data collection, reports, and maintaining relationships with third parties.

Multitasking, time management and excellent interpersonal skills are big priorities for the senior CRA.

According to PayScale , senior clinical research associates earn an average of $98,987.

No matter what stage of your career you are at, it is not too late to consider changing to a remote working position. The amount of people working remotely is increasing as people become more focused on having a good work-life balance. Working remotely allows you the freedom to choose when and where you work without sacrificing pay. By transitioning out of academia and into an industry position, not only will you finally be paid well , but there are many job options that offer the ability to work remotely, including medical writing, scientific editing, consulting, translating, and senior CRA positions. Thanks to the Internet, working from home has never been easier.

If you’re ready to start your transition into industry, you can apply to book a free Transition Call with our founder Isaiah Hankel, PhD or one of our Transition Specialists.  Apply to book a Transition Call here.

Hi, I'm Isaiah Hankel, PhD

I am CEO of Cheeky Scientist, the world's premier career training platform for PhDs. If you want free insights on resumes, LinkedIn, interviewing, careers and more, just enter your details below.

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ABOUT TAVIS MENDEZ, PHD

Tavis Mendez is a scientist in transition who has a passion for benchwork and communicating biomedical research to the public. Dreaming of becoming a scientist since his adolescent years, he obtained his Ph.D. in a parasitology lab, but has also travelled across the USA and to Asia to study cancer and hypertension. In his free time, Tavis loves to conduct culinary experiments in his kitchen.

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• Schools & departments

Biological Sciences PhD

Awards: PhD

Study modes: Full-time, Part-time

Funding opportunities

Programme website: Biological Sciences

Discovery Day

Join us online on 21st August to learn more about postgraduate study at Edinburgh.

Find out more and register

Research profile

Based in King's Buildings Campus at the University of Edinburgh, our staff and students within the School of Biological Sciences are the modern inheritors of a 400-year-old reputation for excellence with Nobel laureates among our notable alumni.

Currently at the leading edge in teaching, research and innovation, we are ranked top 5 in the UK and top 30 in the world for biological sciences in both the QS and Times Higher Education world university rankings (2024).

Key to our success is the highly dynamic and supportive research environment we offer our postgraduate research community; we offer students the tools and confidence they need to make a lasting, positive impact in our world.

Within the School our research is clustered into six large research institutes:

• Institute of Cell Biology
• Institute of Ecology and Evolution
• Institute of Molecular Plant Sciences
• Institute of Immunology & Infection
• Institute for Stem Cell Research (Centre for Regenerative Medicine)
• Institute of Quantitative Biology, Biochemistry and Biotechnology

Join our community and undertake a specialised research project under the guidance of two experienced and well-published supervisors. Our staff have a wide range of research interests and we strongly encourage interdisciplinary research, working closely with physical sciences, engineering, medical and veterinary sciences.

Programme structure

Our PhD programmes are available as full-time or part-time study. The full-time route is 36 months and part-time 72 months, with study normally commencing in October. Both modes of study have an optional additional 12 months, commonly used to write up the final thesis.

The first year of PhD studies is probationary. Your supervisor will identify your training needs and invite you to attend relevant training as appropriate. To successfully progress onto the next year of study students must pass an annual review.

Annual reviews

Students submit a 10-week report and a 10-month report, both assessed by the thesis committee. Each committee comprises of the student's supervisory team, the committee chair, and up to two additional academic staff members.

The 10-week report is a short report outlining the project, with the 10-month report an extended version that includes a literature survey. While meeting their thesis committee, students deliver an oral presentation of their work.

Students complete a presentation during the School’s annual Poster Day. Academic staff and peers review presentations, with feedback discussed with the thesis committee during a 2nd year review meeting.

Students deliver a presentation to academic staff and their peers, traditionally during their Institute’s annual symposium. They also submit a thesis plan for their 3rd year review meeting that contains a timetable for completion, content listings of each chapter, completed work and highlighting outstanding tasks.

To be awarded a PhD, students must submit their final thesis and defend it in an oral examination (assessed according to the University's regulations).

Training and support

We encourage support in a range of academic subjects, transferable skills and personal development. All students who join the programme will have access to our core training and development programme or our specialist cohort-programmes (e.g. EASTBIO).

Our training environment is highly dynamic, with students learning through hands-on, laboratory-based projects as well as guided reading, one-to-one supervision, group seminars, societies and workshops.

The University of Edinburgh Main Library, open to all students, has one of the largest and most important collections in Britain. It has some two million items including the latest publications, key texts, and rare books.

The School of Biological Sciences is based wholly within King's Buildings and in close proximity to the Noreen and Kenneth Murray Library, well stocked with specialist literature, subscription databases and group study rooms and silent study spaces.

Students, dependent on their field of study, will also have access to a range of specialist laboratories and equipment, and state-of-the-art facilities. Examples include:

• imaging facilities including the Central Optical Instrumentation Laboratory (COIL) and Collaborative Optical Spectroscopy Micromanipulation & Imaging Centre (COSMIC)
• advanced flow cytometry facilities, including sorting and analysis
• cutting-edge proteomics facility and automated drug-screening platforms
• Edinburgh Genomics, a University-based sequencing and bioinformatics facility
• multiomics facilities
• the Edinburgh Genome Foundry
• cell culture facilities
• expertise in genomic editing (animal and plant)
• animal and plant facilities

The School also has strong collaborative links to the Royal Botanic Garden Edinburgh to give access to an even broader range of facilities and expertise.

Postgraduate students are given their own desk space in shared student offices and have similar access to facilities available to academic and support staff. Wireless network access is available in many parts of the University campus.

Career opportunities

The School of Biological Sciences offers training events and support for all our postgraduate research students in both academic and key life skills. In collaboration with the University’s Institute of Academic Development (IAD), we also provide a supportive training programme for students hired as tutors and demonstrators.

Entry requirements

These entry requirements are for the 2024/25 academic year and requirements for future academic years may differ. Entry requirements for the 2025/26 academic year will be published on 1 Oct 2024.

A UK 2:1 honours degree, or international equivalent, in a relevant subject. You should have some research experience and be able to demonstrate that you have a good understanding of the field you propose to study.

International qualifications

Check whether your international qualifications meet our general entry requirements:

• Entry requirements by country
• English language requirements

Regardless of your nationality or country of residence, you must demonstrate a level of English language competency at a level that will enable you to succeed in your studies.

English language tests

We accept the following English language qualifications at the grades specified:

• IELTS Academic: total 6.5 with at least 6.0 in each component. We do not accept IELTS One Skill Retake to meet our English language requirements.
• TOEFL-iBT (including Home Edition): total 92 with at least 20 in each component. We do not accept TOEFL MyBest Score to meet our English language requirements.
• C1 Advanced ( CAE ) / C2 Proficiency ( CPE ): total 176 with at least 169 in each component.
• Trinity ISE : ISE II with distinctions in all four components.
• PTE Academic: total 62 with at least 59 in each component.

Your English language qualification must be no more than three and a half years old from the start date of the programme you are applying to study, unless you are using IELTS , TOEFL, Trinity ISE or PTE , in which case it must be no more than two years old.

Degrees taught and assessed in English

We also accept an undergraduate or postgraduate degree that has been taught and assessed in English in a majority English speaking country, as defined by UK Visas and Immigration:

• UKVI list of majority English speaking countries

We also accept a degree that has been taught and assessed in English from a university on our list of approved universities in non-majority English speaking countries (non-MESC).

• Approved universities in non-MESC

If you are not a national of a majority English speaking country, then your degree must be no more than five years old* at the beginning of your programme of study. (*Revised 05 March 2024 to extend degree validity to five years.)

Find out more about our language requirements:

• Academic Technology Approval Scheme

If you are not an EU , EEA or Swiss national, you may need an Academic Technology Approval Scheme clearance certificate in order to study this programme.

Fees and costs

Scholarships and funding

School of Biological Sciences PhD funding

Research scholarships for international students

Edinburgh Doctoral College Scholarship

Search for scholarships and funding opportunities:

• Search for funding

Further information

• Postgraduate Research Administrator
• Phone: +44 (0)131 650 5525
• Contact: [email protected]
• Graduate School of Biological Sciences
• Mary Brück Building
• Colin Maclaurin Road
• The King's Buildings Campus
• Programme: Biological Sciences
• School: Biological Sciences
• College: Science & Engineering

Select your programme and preferred start date to begin your application.

PhD Biological Sciences - 3 Years (Full-time)

Phd biological sciences - 4 years (full-time), phd biological sciences - 8 years (part-time), application deadlines.

We encourage you to apply at least one month prior to entry so that we have enough time to process your application. If you are also applying for funding or will require a visa then we strongly recommend you apply as early as possible, and before the funding deadline.

• How to apply

You must submit two references with your application.

You should submit two references with your application. Before applying you should identify potential supervisors from the School of Biological Sciences and discuss your proposal.

You can find more information on how to apply at our website:

• Postgraduate research at the School of Biological Sciences

Find out more about the general application process for postgraduate programmes:

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6 degrees at 5 universities in the UK.

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Course closed:

Cancer Biology (part-time) is no longer accepting new applications.

The MRes in Cancer Biology (part-time) is offered by the Cancer Research UK Cambridge Centre (CRUK CC) as a part-time course incorporating two long research placements with three short periods out of the lab;  an intense taught course in cancer genomics,  ‘Transferable Skills’ sessions and the third short period offering industry shadowing opportunities. There is also a 36 week course (1 lecture per week) covering general principles of cancer biology as well as insight into how cancers that arise at different sites differ.  

The programme aims to introduce clinicians  and non-clinicians to research skills and specialist knowledge as well as a more general grounding in cancer biology. Its main aims are:

to give students with relevant experience the opportunity to carry out focussed research in the discipline under close supervision; and

to give students the opportunity to acquire or develop skills and expertise relevant to their research interests; and

to give students a broad grounding in both the theory of cancer genomic medicine and also more insight into the practicalities of clinical cancer treatment and care. 

Learning Outcomes

Students should leave the MRes in Cancer Biology (part-time) course with an overview of the basic biology of cancer and of diagnosis, treatment and research frontiers across the range of different tumour types covered by the CRUK Cambridge Centre. Students should also obtain a better understanding of the relationship with industry for the treatment of cancer.

By the end of the MRes programme, students will have:

An understanding of the molecular basis of cancer formation.

Insight into clinical issues related to cancer diagnosis, prognosis and treatment.

Experience of research in two different cancer-related laboratories.

Written two research reports describing their research placements.

Written a PhD proposal that will form the basis of their PhD in the CRUK Cambridge Centre, the expected next destination for the vast majority of this MRes cohort.

The MRes is a part-time 24-month programme that, upon successfully completing to a satisfactory standard, including submission and examination of a suitable PhD proposal, leads directly on to a three-year PhD.

For continuation from the MRes to a probationary PhD, an overall mark of at least 70% is required across the three pieces of work: two research reports and a PhD proposal.  The PhD proposal and oral defence thereof must be at the level of distinction (75%).

The Postgraduate Virtual Open Day usually takes place at the end of October. It’s a great opportunity to ask questions to admissions staff and academics, explore the Colleges virtually, and to find out more about courses, the application process and funding opportunities. Visit the  Postgraduate Open Day  page for more details.

See further the  Postgraduate Admissions Events  pages for other events relating to Postgraduate study, including study fairs, visits and international events.

Key Information

2 years part-time mres + 3 years full-time phd part-time, study mode : research, doctor of philosophy master of research in the first instance, department of oncology, course - related enquiries, application - related enquiries, course on department website, dates and deadlines:, michaelmas 2024 (closed).

Some courses can close early. See the Deadlines page for guidance on when to apply.

Funding Deadlines

These deadlines apply to applications for courses starting in Michaelmas 2024, Lent 2025 and Easter 2025.

Similar Courses

• Cancer Biology PhD
• Medical Science (CRUK CI) MPhil
• Medical Science (CRUK CI) PhD
• Biological Sciences (Crop Science) by advanced study MPhil
• Biological Sciences (Developmental Biology) by advanced study MPhil

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Use your advanced research skills to contribute to new developments in applied biology and biotechnology.

You're viewing program information for local students.

RMIT considers you a local student if you are:

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Research Training Scheme

See admissions

AU$36,480 (2024 annual)

RMIT has a strong reputation for research and teaching in fundamental science and advanced molecular technologies.

Our innovative programs and projects address real-world issues essential to Australia’s innovation agenda.

Research within this discipline uses molecular approaches to diagnose and synthesise solutions for disease and environmental problems.

Under expert supervision, you will develop your skills and join researchers already active in:

• aquatic and marine biology
• ecotoxicology (biomarkers for environmental pollutants)
• bioremediation of contaminated environments
• environmental and molecular approaches to diagnose and develop solutions for disease, agricultural and environmental problems
• genomics and sequencing techniques applied to various organisms
• microbiology (food, medical, environmental and nano biotechnology, vaccine development, fundamentals of host/pathogen interactions)
• plant biology (plant biotechnology and genomics, molecular plant breeding, ecotoxicogenomics, fire and environmental ecology).

RMIT has purpose-built biotechnology labs and advanced microscopy, spectroscopy, tissue culture, microbiology, nanobiotechnology and analytical biochemistry, and experimental animal house facilities located at our Bundoora and City campuses.

How you will learn

Research at rmit, time spent on research.

Full-time candidates are expected to commit at least four days per week (or at least two days per week for part-time candidates) to their research. The academic year is 48 weeks.

Regular contact with your supervisor

A schedule of meetings with your supervisor/s must be established to assess progress against milestones and timely completion.

Resources, facilities and support

You will have access to the Learning Hub and other online and digital resources through the myRMIT student portal.

You will be part of an active research community and have access to resources and workshops to help you succeed.

School of Graduate Research

The School of Graduate Research works with Schools to further support candidates during their postgraduate research degree.

RMIT University is committed to providing you with an education that strongly links formal learning with professional or vocational practice.

We have industry partnerships with organisations and companies such as:  

• Macfarlane Burnet Institute
• Biomass Conversion Technologies
• Bioproperties
• Prince Henry’s Institute
• Department of Agriculture, Fisheries and Forestry.

Learning outcomes

The knowledge and skills you will acquire throughout this degree and how they can be applied in your career are described in the  learning outcomes .

Electives and course plan

You will complete this program under academic supervision.

The PhD program is structured to enable you to:

• complete a compulsory research methods course
• receive training in research integrity and ethics
• select studies in qualitative and quantitative research techniques
• complete a thesis/project which demonstrates your original contribution to the field and your ability to communicate complex or original research for peers and the community to an international standard

Research integrity modules

You are required to complete the online modules:

• Research integrity
• Copyright and intellectual property

Research methods for sciences

Research methods courses step you through the literature review and preparing your research proposal for confirmation of candidature. They are taught in large discipline groups.

You may need to complete an ethics module to ensure your research is ethical and responsible.

Research techniques

You may elect to take (where relevant) electives in qualitative or quantitative research techniques once data collection has begun. You can use your own data to explore different research analysis techniques. Your supervisor will help you decide when you should take these electives.

Co-curricular activities

You are encouraged to participate in activities offered with the university, college and school according to your needs and interests.

This PhD may be undertaken in a project, thesis by publication or thesis mode. Prospective candidates should discuss these modes of submission with their potential supervisor/s.

Course structure

Choose a plan below to find out more about the subjects you will study and the course structure.

*The maximum duration of the PhD program is 4 years full-time and 8 years part-time. However, candidates are expected to complete their program within 3-4 years full-time equivalent and 6-8 years part-time equivalent.

*The maximum duration of the PhD program is 4 years full-time. However, candidates are expected to complete their program within 3-4 years full-time equivalent.

Note: International student visa holders can only study full-time.

You will be able to pursue an academic career in a university or be employed in senior leadership and management positions in government, scientific and industrial research laboratories.

Minimum requirements for admission

Prerequisites, selection tasks.

The minimum requirements for admission to a PhD program are:

• a bachelor degree requiring at least four years of full-time study in a relevant discipline awarded with honours. The degree should include a research component comprised of a thesis, other research projects or research methodology courses that constitute at least 25% of a full-time academic year (or part-time equivalent). The applicant must have achieved at least a distinction average in the final year;  or
• a master degree that includes a research component comprised of at least 25% of a full-time academic year (or part-time equivalent) with an overall distinction average or a master degree without a research component with at least a high distinction average;  or
• evidence of appropriate academic qualifications and/or experience that satisfies the Associate Deputy Vice-Chancellor, Research Training and Development or nominee that the applicant has developed knowledge of the field of study or cognate field and the potential for research sufficient to undertake the proposed program.

At RMIT a grade of distinction represents academic achievement of 70% or higher and a high distinction is 80% or higher.

If you are a current master by research candidate, you are able to apply for a transfer to a doctor of philosophy program through the process prescribed in the  RMIT Higher Degree by Research policy .

There are no prerequisite subjects required for entry into this qualification.

These entrance requirements are the minimum academic standard you must meet in order to be eligible to apply for the program. You will need to complete a selection task as part of your application.

A selection process will be conducted in conjunction with the School and supervisors you nominate.

For further information on the steps you need to take to apply for a research program see  How to apply – Research programs .

English language requirements

Research proposal and supervisor.

You must attach a substantive research proposal that is 2 to 5 pages in length which articulates the intent, significance and originality of the proposed topic using the following headings:

a) title / topic b) research questions to be investigated in the context of existing research/literature in the area c) significance and impact of the research d) methodology / research tasks required to undertake the research e) particular needs (e.g. resources, facilities, fieldwork or equipment that are necessary for your proposed research program, if applicable).

Your application will not be considered if you have not discussed your research topic with a proposed senior and associate supervisor or joint senior supervisors. You must provide the names of the academic staff in the school you have applied to and with whom you have discussed your proposed research.

To study this course you will need to complete one of the following English proficiency tests:

• IELTS (Academic): minimum overall band of 6.5 (with no individual band below 6.0)
• TOEFL (Internet Based Test - IBT): minimum overall score of 79 (with minimum of 13 in Reading, 12 in Listening, 18 in Speaking and 21 in Writing)
• Pearson Test of English (Academic) (PTE (A)): minimum score of 58 (with no communication band less than 50)
• Cambridge English: Advanced (CAE): minimum of 176 with no less than 169 in any component.

For detailed information on English language requirements and other proficiency tests recognised by RMIT, visit  English language requirements and equivalency information .

Don't meet the English language test scores? Complete an English for Academic Purposes (EAP) Advanced Plus Certificate at  RMIT English Worldwide .

You can gain entry to this program from a range of RMIT four year  Bachelor and Honours degrees  or  Postgraduate  or Masters by Research programs.

Fee summary

Fee information for masters by research and doctorate (PhD) programs.

If you are an Australian citizen, Australian permanent resident or New Zealand citizen you may be eligible for a Research Training Scheme (RTS) place where your tuition costs are funded by the Commonwealth Government under the RTS and you have full exemption from tuition fees.

Acceptance in an RTS place is very competitive and places are granted on the condition that you meet annual progress requirements and complete within the allotted time for your program and your status as a part-time or full-time candidate.

This means a maximum of 2 years for a full-time Masters by Research or 4 years for a PhD (or the equivalent part-time).

Contact the School of Graduate Research for more information.

The  student services and amenities fee (SSAF)  is used to maintain and enhance services and amenities that improve your experience as an RMIT student.

In addition to the SSAF there may be  other expenses  associated with your program.

Income tax deductions

Candidates may be eligible to apply for income tax deductions for education expenses linked to their employment. See the  Australian Taxation Office (ATO) website  for more information.

RMIT awards more than 2000 scholarships every year to recognise academic achievement and assist students from a variety of backgrounds.

International applicants

• Fees information  for international candidates looking to study at RMIT's Melbourne campuses.
• PhD  and  masters by research  fees for international candidates studying offshore. 

Other costs

Important fee information.

Find out more details about  how fees are calculated  and the expected annual increase.

Applying for refunds

Find information on how to apply for a  refund  as a continuing international student.

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RMIT University acknowledges the people of the Woi wurrung and Boon wurrung language groups of the eastern Kulin Nation on whose unceded lands we conduct the business of the University. RMIT University respectfully acknowledges their Ancestors and Elders, past and present. RMIT also acknowledges the Traditional Custodians and their Ancestors of the lands and waters across Australia where we conduct our business - Artwork 'Luwaytini' by Mark Cleaver, Palawa.

RMIT University acknowledges the people of the Woi wurrung and Boon wurrung language groups of the eastern Kulin Nation on whose unceded lands we conduct the business of the University. RMIT University respectfully acknowledges their Ancestors and Elders, past and present. RMIT also acknowledges the Traditional Custodians and their Ancestors of the lands and waters across Australia where we conduct our business.

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News Details

Thermo fisher scientific completes acquisition of olink, announces commencement of subsequent offering period.

Subsequent Offering Period Scheduled to Expire at 5:00 p.m., New York Time, on July 16, 2024

WALTHAM, Mass.--(BUSINESS WIRE)-- Thermo Fisher Scientific Inc. (NYSE: TMO) (“Thermo Fisher”), the world leader in serving science, today announced that it has completed its acquisition of Olink Holding AB (publ) (“Olink”) (Nasdaq: OLK), a leading provider of next-generation proteomics solutions. The transaction values Olink at approximately $3.1 billion, net of $96 million of acquired cash. Olink will become part of Thermo Fisher's Life Sciences Solutions segment.

“We are excited to welcome Olink colleagues to Thermo Fisher,” said Marc N. Casper, chairman, president and chief executive officer of Thermo Fisher. “Protein research is key to advancing our understanding of human biology and delivering on the promise of precision medicine. Olink’s proven and transformative solutions for proteomics are highly complementary to our leading mass spectrometry and life sciences offerings. By expanding our capabilities in the high-growth proteomics market, we will enable scientists and researchers around the globe to meaningfully accelerate discovery and scientific breakthroughs while delivering significant value to our shareholders.”

The Offer was made pursuant to the Purchase Agreement, dated as of October 17, 2023, by and between Thermo Fisher and Olink and as described in the tender offer statement on Schedule TO, as amended and supplemented from time to time, filed by Thermo Fisher with the U.S. Securities and Exchange Commission (the “SEC”) on October 31, 2023 (the “Tender Offer Statement”). Thermo Fisher, through its wholly owned subsidiary, Orion Acquisition AB (the “Buyer”), today has accepted for purchase all of the outstanding common shares of Olink (“Shares”) and all of the outstanding American Depositary Shares, each representing one Share of Olink (“ADSs” and collectively with the Shares, “Offer Securities”) that were validly tendered and not properly withdrawn prior to the expiration time of the initial offering period for the Offer Securities for $26.00 per Share and per ADS, in cash (the “Offer”). Buyer will promptly pay cash for tendered and accepted Offer Securities. Upon settlement of the Offer, Thermo Fisher will own Offer Securities representing approximately 98.2% of Olink’s outstanding Shares.

Further Transaction Details and Timing

The initial offering period for the Offer and withdrawal rights expired as scheduled at 5:00 p.m., New York time, on July 9, 2024 (the “Expiration Time”). DNB Markets, a part of DNB Bank ASA, Sweden Branch, the depositary and paying agent for the Offer with respect to the Shares, has advised Thermo Fisher and Buyer that, as of the Expiration Time, approximately 84,755,626 Shares have been validly tendered and not properly withdrawn pursuant to the Offer. The Bank of New York Mellon, the tender agent for the Offer with respect to the ADSs, has advised Thermo Fisher and Buyer that, as of the Expiration Time, approximately 37,676,600 ADSs have been validly tendered and not properly withdrawn pursuant to the Offer. These Shares and ADSs collectively represent approximately 98.2% of the outstanding Shares. With all conditions to the Offer satisfied, on July 10, 2024, Buyer has accepted for purchase all Offer Securities that were validly tendered and not properly withdrawn pursuant to the Offer as of the Expiration Time, and payment for such Offer Securities will be made promptly in accordance with the terms of the Offer.

Thermo Fisher also announced the commencement of a subsequent offering period (the “Subsequent Offer”) scheduled to expire at 5:00 p.m., New York time, on July 16, 2024, unless extended (the “Subsequent Offering Period”), as more fully described in the Tender Offer Statement. All Shares and ADSs validly tendered during the Subsequent Offering Period will be immediately accepted for payment, and tendering holders will thereafter promptly be paid the same Offer price of $26.00 per Share or per ADS, as applicable, in cash, without interest and less applicable withholding taxes or other taxes, upon the terms and subject to the conditions set forth in the Offer to Purchase dated as of October 31, 2023 (together with any amendments and supplements thereto, the “Offer to Purchase”). The procedures for accepting the Offer and tendering Offer Securities during the Subsequent Offering Period are the same as those applicable to the Offer as described in the Offer to Purchase, except that Shares and ADSs validly tendered during the Subsequent Offering Period may not be withdrawn. To acquire any Shares that remain untendered following the completion of the Subsequent Offer, Thermo Fisher intends to initiate a compulsory redemption under the Swedish Companies Act (Sw. aktiebolagslagen (2005:551)).

Olink holders who have validly tendered and not properly withdrawn their Shares or ADSs of Olink prior to the Expiration Time do not need to re-tender their Shares or ADSs or take any other action as a result of the Subsequent Offer.

Delisting & Deregistration of Olink’s ADSs

Olink has requested that the NASDAQ Global Market (“Nasdaq”) suspend trading of Olink ADSs and file a Form 25, Notification of Removal from Listing and/or Registration with the SEC, thereby commencing the process of delisting and deregistering the ADSs. Provided that the criteria for deregistration are met, Olink also intends to terminate its reporting obligations under the Securities Exchange Act of 1934, as amended, by filing a Certification and Notice of Termination of Registration on Form 15 with the SEC. Following delisting from Nasdaq, ADSs of Olink will not be listed or registered on another national securities exchange. Delisting is likely to reduce significantly the liquidity and marketability of any ADSs of Olink that have not been tendered pursuant to the Offer or the Subsequent Offer.

Requests for copies of the Offer to Purchase (as amended or supplemented from time to time), the ADS letter of transmittal and the acceptance form for Shares (including the instructions attached thereto) and other Offer materials may be directed to the call service of Georgeson LLC at +1-866-821-2550 (U.S. toll-free), +1-781-222-0033 (outside U.S. & Canada) or +46-846-007-389 (Sweden), or via email at [email protected] . A copy of these documents may be obtained at the website maintained by the SEC at www.sec.gov .

For Thermo Fisher, Cravath Swaine & Moore LLP and Advokatfirman Vinge KB served as legal counsel. For Olink, J.P. Morgan Securities LLC served as lead financial advisor, Goldman Sachs Bank Europe SE, Sweden Bankfilial served as financial advisor and Baker & McKenzie served as legal counsel.

About Thermo Fisher Scientific

Thermo Fisher Scientific Inc. is the world leader in serving science, with annual revenue over $40 billion. Our Mission is to enable our customers to make the world healthier, cleaner and safer. Whether our customers are accelerating life sciences research, solving complex analytical challenges, increasing productivity in their laboratories, improving patient health through diagnostics or the development and manufacture of life-changing therapies, we are here to support them. Our global team delivers an unrivaled combination of innovative technologies, purchasing convenience and pharmaceutical services through our industry-leading brands, including Thermo Scientific, Applied Biosystems, Invitrogen, Fisher Scientific, Unity Lab Services, Patheon and PPD. For more information, please visit www.thermofisher.com .

Forward-looking Statements

This press release contains forward-looking statements that involve a number of risks and uncertainties. Words such as “believes,” “anticipates,” “plans,” “expects,” “seeks,” “estimates,” and similar expressions are intended to identify forward-looking statements, but other statements that are not historical facts may also be deemed to be forward-looking statements. Important factors that could cause actual results to differ materially from those indicated by forward-looking statements include risks and uncertainties relating to: the COVID-19 pandemic, the need to develop new products and adapt to significant technological change; implementation of strategies for improving growth; general economic conditions and related uncertainties; dependence on customers’ capital spending policies and government funding policies; the effect of economic and political conditions and exchange rate fluctuations on international operations; use and protection of intellectual property; the effect of changes in governmental regulations; any natural disaster, public health crisis or other catastrophic event; and the effect of laws and regulations governing government contracts, as well as the possibility that expected benefits related to recent or pending acquisitions may not materialize as expected; Olink’s business experiencing disruptions due to transaction-related uncertainty or other factors making it more difficult to maintain relationships with employees, customers, licensees, other business partners or governmental entities; difficulty retaining key employees; the outcome of any legal proceedings related to the compulsory redemption; and the parties being unable to successfully implement integration strategies or to achieve expected synergies and operating efficiencies within the expected time-frames or at all. Additional important factors that could cause actual results to differ materially from those indicated by such forward-looking statements are set forth in Thermo Fisher’s Annual Report on Form 10-K and subsequent quarterly report on Form 10-Q, which are on file with the U.S. Securities and Exchange Commission (“SEC”) and available in the “Investors” section of Thermo Fisher’s website, ir.thermofisher.com, under the heading “SEC Filings”, and in any subsequent documents Thermo Fisher files or furnishes with the SEC, and in Olink’s Annual Report on Form 20-F and subsequent interim report on Form 6-K, which are on file with the SEC and available in the “Investor Relations” section of Olink’s website, https://investors.olink.com/investor-relations , under the heading “SEC Filings”, and in any subsequent documents Olink files or furnishes with the SEC. While Thermo Fisher or Olink may elect to update forward-looking statements at some point in the future, Thermo Fisher and Olink specifically disclaim any obligation to do so, even if estimates change and, therefore, you should not rely on these forward-looking statements as representing either Thermo Fisher’s or Olink’s views as of any date subsequent to today.

Additional Information and Where to Find It

This communication is for informational purposes only and is neither an offer to purchase nor a solicitation of an offer to sell any common shares or American Depositary Shares of Olink or any other securities, nor is it a substitute for the tender offer materials that Thermo Fisher or the Buyer has filed with the SEC. The terms and conditions of the tender offer are published in, and the offer to purchase common shares and American Depositary Shares of Olink is made only pursuant to, the offer document and related offer materials prepared by Thermo Fisher and the Buyer and is filed with the SEC in a tender offer statement on Schedule TO. In addition, Olink has filed a solicitation/recommendation statement on Schedule 14D-9 with the SEC with respect to the tender offer.

THE TENDER OFFER MATERIALS (INCLUDING AN OFFER TO PURCHASE, AN ADS LETTER OF TRANSMITTAL, ACCEPTANCE FORM FOR SHARES AND CERTAIN OTHER TENDER OFFER DOCUMENTS) AND THE SOLICITATION/RECOMMENDATION STATEMENT ON SCHEDULE 14D-9, AS THEY MAY BE AMENDED FROM TIME TO TIME, CONTAIN IMPORTANT INFORMATION. INVESTORS AND SHAREHOLDERS OF OLINK ARE URGED TO READ THESE DOCUMENTS CAREFULLY BECAUSE THEY, AND NOT THIS DOCUMENT, GOVERN THE TERMS AND CONDITIONS OF THE TENDER OFFER, AND BECAUSE THEY CONTAIN IMPORTANT INFORMATION THAT SUCH PERSONS SHOULD CONSIDER BEFORE MAKING ANY DECISION REGARDING TENDERING THEIR COMMON SHARES AND AMERICAN DEPOSITARY SHARES.

The tender offer materials, including the offer to purchase, the related ADS letter of transmittal and acceptance form for shares and certain other tender offer documents, and the solicitation/recommendation statement and other documents filed with the SEC by Thermo Fisher or Olink, may be obtained free of charge at the SEC's website at www.sec.gov , at Olink’s website https://investors.olink.com/investor-relations , at Thermo Fisher's website at www.thermofisher.com or by contacting Thermo Fisher's investor relations department at 781-622-1111. In addition, Thermo Fisher's tender offer statement and other documents it will file with the SEC will be available at https://ir.thermofisher.com/investors .

Media Contact Information: Sandy Pound Phone: 781-622-1223 E-mail: [email protected] Website: www.thermofisher.com

Investor Contact Information: Rafael Tejada Phone: 781-622-1356 E-mail: [email protected]

Investor Relation Kit

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Rafael Tejada

Vice President, Investor Relations

Thermo Fisher Scientific

168 Third Avenue

Waltham, MA USA 02451

781.622.1111 E-mail: [email protected]

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Email Alert Sign Up Confirmation Thermo Fisher Scientific Inc. (NYSE: TMO) is the world leader in serving science, with annual revenue exceeding $30 billion. Our Mission is to enable our customers to make the world healthier, cleaner and safer. Whether our customers are accelerating life sciences research, solving complex analytical challenges, improving patient diagnostics and therapies or increasing productivity in their laboratories, we are here to support them. Our global team of more than 80,000 colleagues delivers an unrivaled combination of innovative technologies, purchasing convenience and pharmaceutical services through our industry-leading brands, including Thermo Scientific, Applied Biosystems, Invitrogen, Fisher Scientific, Unity Lab Services and Patheon. For more information, please visit www.thermofisher.com. Thermo Fisher Scientific Inc. is the world leader in serving science, with annual revenue of more than $40 billion. Our Mission is to enable our customers to make the world healthier, cleaner and safer. Whether our customers are accelerating life sciences research, solving complex analytical challenges, increasing productivity in their laboratories, improving patient health through diagnostics or the development and manufacture of life-changing therapies, we are here to support them. Our global team delivers an unrivaled combination of innovative technologies, purchasing convenience and pharmaceutical services through our industry-leading brands, including Thermo Scientific, Applied Biosystems, Invitrogen, Fisher Scientific, Unity Lab Services, Patheon and PPD. For more information, please visit www.thermofisher.com. Thermo Fisher Scientific Inc. is the world leader in serving science, with annual revenue over $40 billion. Our Mission is to enable our customers to make the world healthier, cleaner and safer. Whether our customers are accelerating life sciences research, solving complex analytical challenges, increasing productivity in their laboratories, improving patient health through diagnostics or the development and manufacture of life-changing therapies, we are here to support them. Our global team delivers an unrivaled combination of innovative technologies, purchasing convenience and pharmaceutical services through our industry-leading brands, including Thermo Scientific, Applied Biosystems, Invitrogen, Fisher Scientific, Unity Lab Services, Patheon and PPD. For more information, please visit  www.thermofisher.com . Corporate Social Responsibility Information Security Thermo Scientific Applied Biosystems Fisher Scientific Unity Lab Services follow us on facebook follow us on twitter follow us on linkedin follow us on youtube follow us on instagram © Copyright Thermo Fisher Scientific Inc. 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