See the website for a list of electives: http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements
The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2024 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.
Code | Title | Course Units |
---|---|---|
Year 1 | ||
Fall | ||
Advanced Topics in Current Biological Research | ||
Cell Biology | ||
Advanced Evolution | ||
Independent Study and Research | ||
Spring | ||
Genetic Analysis | ||
Evolutionary Ecology | ||
Theoretical Population Biology | ||
Independent Study and Research | ||
Independent Study and Research | ||
Summer | ||
Year 2 | ||
Fall | ||
Independent Study and Research | ||
Spring | ||
Communication for Biologists | ||
Independent Study and Research | ||
Summer | ||
Year 3 and Beyond | ||
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On this page:, at a glance: program details.
Degree Awarded: PHD Evolutionary Biology
The evolutionary biology PhD is a transdisciplinary graduate degree program that provides doctorate-level training in the historical, conceptual, empirical and quantitative aspects of biological evolution.
Evolution is a fundamental scientific concept underlying all aspects of modern biological, environmental and health-related research. It cuts across biological sciences in ways that few other foci do and informs the theoretical foundations of subfields like population genetics and ecosystem ecology. It allows integration of information and patterns across levels of organization, informs the theoretical foundations of subfields ranging from population genetics to systematics to ecosystem ecology, and provides bridges between temporal and spatial scales.
Forgoing emphases on particular taxa or methods, the program focuses on understanding the patterns and processes that have shaped life on Earth and continue to do so, training the next generation of scientists to use this knowledge to meet present and future challenges to the biosphere and human health in the face of increasing environmental perturbation.
Program Faculty PhD Students
How to apply.
Applications open September 1 for admission in Fall of the following year. The application deadline is December 1 . We accept applications for Fall semesters only. We cannot guarantee that applications received after the December 1 deadline will be considered for admission.
All applicants must apply by filling out ASU's Graduate Admissions application. All application materials must be submitted through the application or to Graduate Admissions directly. Please do not mail or email any documents to the School of Life Sciences.
Required materials and information include the following:
Following the December 1 deadline, faculty will begin reviewing applications. Applicants should monitor their My ASU priority tasks to ensure there are no missing materials in their application.
Faculty will decide which applicants they would like to invite to our Graduate Recruitment Weekends (GRWs), typically held in February. Applicants will hear from the School of Life Sciences in January if they are invited to participate in the GRWs.
Admission decisions will begin after the GRWs, and applicants typically receive final decisions by April 1.
Minimum requirements for admission include the following:
Desired qualifications typically seen in competitive candidates:
Please note that the GRE is not required.
Students offered admission to a PhD program in the School of Life Sciences will typically receive a funding offer as well. While individual funding offers may differ to some degree, they typically include teaching assistant and/or research assistant positions each semester (summer optional) for 5 years. These positions provide financial coverage through the following:
To discover more, check out the ASU Graduate College's funding opportunities !
84 credit hours, a written comprehensive exam, an oral comprehensive exam, a prospectus and a dissertation
Required Core (9 credit hours) BIO 514 Statistical Models for Biology (4) EVO 601 Principles of Evolution (3) EVO 610 Research Areas of Evolution (2)
Electives (9 credit hours) The program advisor as well as the student's advisor will determine these courses in conjunction with the student.
Other Requirements (54 credit hours) research, coursework or 30 credit hours from a previously awarded master's degree
Dissertation (12 credit hours) EVO 799 Dissertation (12)
Additional Curriculum Information Students take EVO 610 twice for one credit hour.
Applicants must fulfill the requirements of both the Graduate College and The College of Liberal Arts and Sciences.
Applicants are eligible to apply to the program if they have earned a bachelor's or master's degree in a related discipline from a regionally accredited institution.
Applicants must have a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in the last 60 hours of their first bachelor's degree program, or a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in an applicable master's degree program.
Applicants must submit the following:
Additional Application Information An applicant whose native language is not English must provide proof of English proficiency regardless of their current residency.
Learn about our programs, apply to a program, visit our campus, learning outcomes.
Those who have earned a doctorate in evolutionary biology are prepared for academic careers at every level, from community colleges to research universities, and their skills and knowledge are also valuable for government careers in federal and state agencies, and for careers in industry and nongovernmental organizations.
Career examples include:
If you have questions related to admission, please click here to request information and an admission specialist will reach out to you directly. For questions regarding faculty or courses, please use the contact information below.
INFORMATION FOR
Associate Professor of Molecular Biophysics and Biochemistry, and of Cell Biology
Assistant Professor
Associate Professor of Molecular, Cellular and Developmental Biology and of Physics and of Neuroscience
William R. Kenan, Jr. Professor of Molecular Biophysics and Biochemistry; Head, Branford College
Lewis B. Cullman Professor of Molecular, Cellular and Developmental Biology and Professor of Physics
Fergus F. Wallace Professor of Genetics
Sharon Hammes-Schiffer View Full Profile
Professor of Molecular, Cellular and Developmental Biology
Eugene Higgins Professor of Molecular Biophysics and Biochemistry and Professor of Physics
Associate Professor of Pharmacology
Associate Professor of Molecular Biophysics and Biochemistry
Professor; Assoc Prof Dept of Mechanical Engineering & Materials Science and Physics; Associate Professor
Assistant Professor of Molecular, Cellular and Developmental Biology
Associate Professor of Genetics and Cell Biology
Assistant Professor, Molecular, Cellular and Developmental Biology
Assistant Professor of Molecular Biophysics and Biochemistry
Professor of Cell Biology
Publications, coordinates.
The Department of Theoretical Biology uses mathematics and computer simulations to study the dynamics of evolution. Our team is interdisciplinary, with backgrounds ranging from biology, physics, and mathematics to computer science and economics. Our goal is, on the one hand, to model biological systems with a close connection to empirical research, on the other hand we develop the mathematical toolbox of theoretical biology further and aim to generate new conceptual insights. The biological problems we address include the emergence and maintenance of polymorphisms in natural populations, the evolution of cooperation and the evolutionary dynamics of cancer.
The department is organised into the three main themes Cancer Evolution, Metaorganisms, and Population Structure and Game Theory. In addition, the department is hosting four research groups, which independently pursue their own research directions.
The accumulation of mutations within the cells of a tissue can be viewed as an evolutionary process. We use mathematical and computational models to gain insights into the dynamics of these processes. In collaboration with clinicians, experimentalists and bioinformaticians, we have so far mostly focussed on blood diseases, but several general insights can be used to construct theoretical models for other systems.
Why do individuals from different taxonomic groups and even kingdoms of life interact with each other within a metaorganism in a generally beneficial way and thereby form a unit of selection? This question is one of the most fascinating, unresolved mysteries in current evolutionary biology. We develop mathematical models to understand interactions within the metaorganism in close collaboration with the CRC 1182 Origin and Function of Metaorganisms .
While the evolution of cooperation is the most popular topic in evolutionary game theory, the game theoretical approach offers much more to evolutionary biology. We are interested in eco-evoluionary dynamics under frequency and density dependent selection, in the evolution of multicellular units, but also in the general role that population structure (in terms of groups, networks or age) plays for evolutionary dynamics.
In the Microbial Evolutionary Dynamics group, we use bacterial and phage populations to empirically investigate fundamental evolutionary processes. Our over-arching theme is the evolution and maintenance of genetic structure – how and why particular genetic entities come to exist.
Research group theoretical models of eco-evolutionary dynamics.
Evolution proceeding at ecological time scales changes the way we approach traditional evolutionary dynamics. We are interested in understanding the evolutionary change in interactions in the light of the ecology in which they take place.
Chaitanya is now a Professor at the University of Würzburg, but will maintain his group at the Max Planck Institute for Evolutionary Biology.
Research group stochastic evolutionary dynamics.
Our research lies at the intersections of evolution with ecology and with medicine. Using mathematical models, we study which factors promote or hamper rapid adaptation to changing environments.
Research group dynamics of microbial collectives.
Microbial ecology often deploys in the context of collectives -- communities, biofilms, multicellular aggregates. We inquire how the properties of such collectives emerge and evolve based on cellular-level mechanisms.
Research group molecular systems evolution.
Our research aims at understanding the evolution of genomes. By comparing genome sequences from various individuals and/or species, one can gain insights into the process of how and when species are created. We combine modeling approaches and statistical data analysis, as well as lab experiments when possible.
Prof. dr. arne traulsen.
Group leader
Dr. philipp altrock.
Project Leader
Dr. ernesto berríos-caro.
Postdoc (Uecker group)
PhD candidate (Uecker group)
PhD candidate (Dutheil group)
Małgorzata fic.
PhD candidate (Gokhale group)
Finote gijsman.
Visiting PhD candidate
Master's student (Dutheil group)
Jinyang liang, sungbin lim.
PhD candidate (Gallie group)
Dr. emil mallmin, guy ngan wing yui, dharanish rajendra, maría alejandra ramírez.
PhD candidate
Dr. nikhil sharma, dr. michael sieber.
Staff scientist
Qianci yang, general advice.
We are an interdisciplinary department and people working here have backgrounds ranging from theoretical physics and computer science to biology and medicine. The combination of different skillsets is crucial for the success of our work. A central aspect in this is that applicants are highly motivated to do research in our department. Thus, please let us know in your application why our department seems to be the right place for you and why you are the right person for us! How did you hear about us? Which papers did you read? Did someone inspire you? Tell us about the path that led you to your interest in the department! Do you already have a concrete project in mind that you would like to work on? Or are you interested in the general area? In the end, there must be the right match on both sides - you need to be sure that this is the place to develop your career further and we need to be sure that you will fit in scientifically and as a person!
If you apply for a job at the MPI Plön, please send
There are two tracks to join our department:
PNAS 121:e2312822121 (2024)
E. Mallmin, A. Traulsen and S. De Monte
PNAS 120:e2311584120 (2023)
C. M. Saad-Roy and A. Traulsen
Evolution 77:1408-1421 (2023)
M. Raatz and A. Traulsen
Journal of the Royal Society Interface 20:20220769 (2023)
S. Yagoobi, N. Sharma and A. Traulsen
© 2023 Department of Theoretical Biology
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Theoretical biology is a subdiscipline of biology that may have a significant impact on the further development of biology. Its meaning for the biological sciences is comparable to the meaning of theoretical physics for the physical sciences. Theoretical physics, and especially mathematical physics, is concerned with the description of physical phenomena in mathematical terms. Beyond this, however, it makes abstractions from physical reality and puts them into a conceptual framework in such a way that from abstract models predictions can be made about the phenomenology of real systems. Theoretical biology is trying to do just that; and the fact that in doing so many concepts of the physical sciences are “taken over” and used to describe biological phenomena in an abstract manner provides an argument for considering theoretical biology as a legitimate domain in biophysics. Its aim, of course, is the unification of assemblies of facts and their simplification through the logic of a (abstract) concept, thus arriving at a deeper understanding and greater predictability.
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© 1989 Kluwer Academic Publishers
Sybesma, C. (1989). Theoretical biology. In: Biophysics. Springer, Dordrecht. https://doi.org/10.1007/978-94-009-2239-6_12
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Prof. Jan Pieter Abrahams Uncovering the fundamental shapes of life
Prof. Attila Becskei Mechanisms controlling gene regulatory networks
Prof. Flavio Donato Development of neuronal circuits and cognitive functions
Prof. Knut Drescher Development and functions of bacterial communities
Prof. Ben Engel Charting the molecular architecture of organelles
Prof. Stephan Grzesiek Insights into the structure and function of biomolecules
Prof. Richard Neher Microbial evolutions and adaptation
Prof. Torsten Schwede Computational modeling of 3D protein structures
Prof. Erik van Nimwegen Function and evolution of genome-wide regulatory networks
Prof. Mihaela Zavolan Key regulators of gene expression and cell identity
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Phd research at the interface of quantitative science and biology.
HMS Campus, 240 Longwood Avenue, Seeley G. Mudd Bldg., Room 204c Boston, MA 02115
Our lab studies microbial evolution, primarily of antibiotic resistance, with a goal of developing practical interventions to reduce or reverse resistance.
... Read more about Michael Baym, Ph.D. (He/ Him/ His)
Transcription factors and DNA regulatory elements.... Read more about Martha L. Bulyk, Ph.D. (She/ Her/ Hers)
Pursuing "Next generation" DNA sequencing methods and chip-DNA-synthesis, gene editing and stem cell engineering. ... Read more about George M. Church, Ph.D. (He/ Him/ His)
Physical tools to study molecules, cells, and organisms.... Read more about Adam E. Cohen, Ph.D. (He/ Him/ His)
We study the neural circuit and molecular basis of individual differences in behavior using custom high-throughput instrumentation, with an eye to its ultimate cause in evolution. ... Read more about Benjamin de Bivort
Molecular Principles Enabling Structural Diversification of Very Long-Chain Fatty Acids.... Read more about Vladimir Denic
We use both theory and experiments to study evolutionary dynamics and population genetics, particularly in situations where natural selection is pervasive.... Read more about Michael M. Desai, Ph.D.
Statistical computations and neuronal mechanisms underlying complex decisions and behavior under uncertainty. The work in the lab is theoretical in nature, and we collaborate with experimentalists in a close loop to refine both theories and experiments.... Read more about Jan Drugowitsch, Ph.D. (He/ Him/ His)
Neural circuits and behavior in larval zebrafish... Read more about Florian Engert
The Farnung Lab is interested in molecular mechanisms at the interface of transcription and chromatin.... Read more about Lucas Farnung
Structural biology of signaling and transport through biological membranes.... Read more about Rachelle Gaudet, Ph.D. (She/ Her/ Hers)
Our laboratory investigates the molecular mechanisms of the vertebrate immune system.
... Read more about Sun Hur, Ph.D. (She/ Her/ Hers)
Chromosome Mechanics and Dynamics... Read more about Nancy Kleckner, Ph.D.
Biophysics is an interdisciplinary science that applies the approaches and methods of physics to study biological systems . Biophysics covers all scales of biological organization , from molecular to organismic and populations.
Daniel fisher | physicists and evolution: puzzles and expectations.
To most physicists, almost everything about evolution is puzzling. What determines how…
To most physicists, almost everything about evolution is puzzling. What determines how much complexity can — and has — evolved? Why is there so much biological diversity on all scales of differences? How could evolution…
The CLS PhD program is organized by the Theoretical Biology and Bioinformatics group of the Institute for Biodynamics and Biocomplexity performing interdisciplinary research that combines biology, physics, chemistry and mathematics to understand living systems at different organization levels from individual molecules, such as DNA, lipids and proteins, to cells and model organisms. Through close interaction between experimentation, theory and simulation, we aim to achieve a systems-level understanding of complex dynamic processes and their evolution. One possible route into this program is via the graduate program Quantitative Biology , which allows the best students to write their own PhD proposal.
Current expertise in the Computational Life Sciences PhD program involves a variety of biological disciplines and computational approaches. Biological areas that we cover are genome evolution, eco-evolutionary dynamics, gene regulatory networks, immunology, cell motility, development, and spatial pattern formation. This enables the PhD program to train students in Systems Biology in a variety of biological backgrounds. We cover computational approaches ranging from large scale bioinformatic data analysis, varying from genomic to gene expression data, mathematical analysis, to computational modeling, where the latter varies from the Cellular Potts model that is used to study development, evolutionary computation, to discrete-event agent based computer simulation models.
The main objective of the Computational Life Sciences PhD program is to train students to become excellent and independent researchers in Theoretical Biology and Bioinformatics, with solid expertise in modeling and bioinformatics, and a good interdisciplinary knowledge in the life sciences. Specifically, students should be able to
The Computational Life Sciences PhD program is part of the Graduate School of Life Sciences and the Life Sciences and Biocomplexity focus area of Utrecht University. The director of the program is Prof. Dr. Rob J. de Boer . All staff members of the Theoretical Biology and Bioinformatics group in this focus area will supervise PhD students, give courses, run journal clubs, and international seminars.
In the past we have attracted PhD students from all over the world and with various scientific backgrounds. The basic requirement for admission is a relevant and sufficient scientific background in Computational Life Sciences, and financial support to pay for the PhD position. Because of the interdisciplinary nature of our PhD program the admission criteria will have to be flexible and individualized. The track Theoretical Biology and Bioinformatics in the master program Biology and Biocomplexity of the Graduate School of Life Sciences provides an optimal preparation for a PhD in Computational Life Sciences. Students with a master education in mathematics or physics can also be admitted by taking additional courses in the relevant biological areas and/or modeling and bioinformatics. Similarly, students with a biological master, but an insufficient background in modeling and bioinformatics, will have to repair these computational skills to be admitted to the school. An excellent course for repairing deficiencies in background is the MSc course ``Bioinformatic Processes" that is mandatory for all MSc students in our MSc track Theoretical Biology and Bioinformatics.
PhD students will be supervised by at least one of the staff members of the Theoretical Biology and Bioinformatics group. We recommentd the PhD student and the supervising staff member to formulate an appropriate Course Plan together. The contents of the educational program, and the quality of the program, i.e., the scientific level of the courses the PhD student will attend, are the responsibility of the supervising staff member and the program coordinator. After the PhD has been completed, any PhD student with 20 ECTs is eligible for a Graduate School of Life Sciences certificate.
PhD students in our group attend the advanced courses that we give ourselves, and various other national and international courses. Examples of schools and courses attended by our PhD students:
In Holland PhD positions come with a salary. This means that the group has to have funding to hire you as a PhD student. Check our webpage Positions to check whether we have open PhD positions available. Note that we always require a solid background in Theoretical Biology and/or Bioinformatics and a genuine interest in biology. Theoretical Biology & Bioinformatics / Last modified on 20 July 2020 / Webmaster: R.J.DeBoer@uu.nl
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The Center for Theoretical Biological Physics (CTBP) at Rice University is one of 10 Physics Frontier Centers supported by the National Science Foundation through the Physics Division of the Directorate for Mathematical and Physical Sciences, with additional sponsorship from three NSF divisions: Molecular and Cell Biology, Chemistry, and Materials Research. The NSF established the CTBP to facilitate research and support education in biological physics, a rapidly emerging area of science that uses concepts and methodologies from biology, math, and physics to better understand the complexity of living systems.
Originally headquartered at the University of California-San Diego (UCSD), the CTBP moved to Rice in 2011 upon recruitment of biophysicists José Onuchic and Peter Wolynes and bioengineer Herbert Levine with funding from the Cancer Prevention & Research Institute of Texas (CPRIT). The Center is now located at Rice’s BioScience Research Collaborative , and while some of its research efforts are still conducted at UCSD (TMC news), it has formed additional partnerships with the Baylor College of Medicine, the University of Houston, and The University of Texas Health Science Center – Houston, all of which are close to the Rice campus.
Taking advantage of its sophisticated computational and experimental facilities, CTBP researchers work on four broadly defined areas:
Besides research, the CTBP is committed to education and outreach. The Center offers many research opportunities to postdoctoral fellows and graduate and undergraduate students. These Opportunities for Research in Biophysics, Informatics and Theoretical Science (ORBITS), encourage using both theoretical and experimental physics to further the understanding of biology and its application to biomedicine.
In addition, the CTBP is the lead institute for Physics of Living Systems, a novel network community where graduate students can help each other navigate the difficulties in dealing with research in biological physics, an inherently multi-disciplinary field that is still relatively new in many institutions.
For outreach, the CTBP partners with the University of Houston and the Houston Community College to establish programs to enable undergraduates from under-represented minorities to experience cutting-edge research, thereby encouraging them to consider attending graduate school in a STEM field.
https://ctbp.rice.edu/
http://www.tmc.edu/news/2014/09/nsf-renews-grant-for-biological-physics-research-at-rice/
December 2018.
Ottawa-Carleton Joint Program
Established in 1984, the Ottawa-Carleton Institute of Biology (OCIB) combines the research strengths of the University of Ottawa and Carleton University. The Institute offers graduate programs leading to the master’s (MSc) and doctoral (PhD) degrees in Biology.
Research facilities are shared between the two campuses. Students have access to the professors, courses and facilities at both universities.
The Institute is a participating unit in the collaborative program in Chemical and Environmental Toxicology (at the master’s and doctoral levels).
The doctoral program participates in the Combined Program for Degrees in Medicine and Philosophy (MD/PhD). For more information please see the website of the Faculty of Medicine .
The estimated amount for university fees associated with this program are available under the section Finance your studies .
International students enrolled in a French-language program of study may be eligible for a differential tuition fee exemption .
Graduate studies office, faculty of science, 30 marie-curie street, gendron hall, room 181, ottawa, ontario, canada, tel.: 613-562-5800 x 3145, email: [email protected] , twitter | faculty of science, facebook | faculty of science.
For the most accurate and up to date information on application deadlines, language tests and other admission requirements, please visit the specific requirements webpage.
Note: International candidates must check the admission equivalencies for the diploma they received in their country of origin.
Note: International students must provide proof of financial support: i.e., a stipend provided by a supervisor as well as a combination of awards and/or trust funds.
Applicants must be able to understand and fluently speak the language of instruction (French or English) in the program to which they are applying. Proof of linguistic proficiency may be required.
Applicants whose first language is neither French nor English must provide proof of proficiency in the language of instruction.
Note: Candidates are responsible for any fees associated with the language tests.
Students enrolled in the master’s program in biology at the University of Ottawa may be eligible to fast-track directly into the doctoral program without writing a master’s thesis, provided the following conditions are met:
Note: If the student meets the requirements, the transfer must take place within sixteen months of initial enrollment in the master’s. The minimal admission average requirements for the doctoral program must also be met. Following transfer, all the requirements of the doctoral program must be met.
Requirements for this program have been modified. Please consult the 2022-2023 calendars for the previous requirements.
The Department may require students to take additional courses depending on their backgrounds.
Students must meet the following requirements:
Code | Title | Units |
---|---|---|
Compulsory Courses | ||
Research and Communication in Biology | 3 Units | |
3 optional course units in biology (BIO) at the graduate level | 3 Units | |
Comprehensive Examination | ||
Comprehensive Examination | ||
Thesis | ||
Doctoral Thesis |
BIO 8950 is replaced with an additional 3 optional course units in biology (BIO) if previously taken BIO 5950 .
The optional course units may also be selected in related disciplines approved by the Department of Biology.
The comprehensive examination must be completed within twelve months of the initial admission into the program.
Presentation and successful defence of a thesis based on original research carried out under the direct supervision of a faculty member of the Department.
Students are responsible for ensuring they have met all of the thesis requirements .
The passing grade in all courses is B.
Students who fail 6 units, or the thesis proposal, or the comprehensive exam, or whose progress is deemed unsatisfactory must withdraw from the program.
Located in the heart of Canada’s capital, a few steps away from Parliament Hill, the University of Ottawa is among Canada’s top 10 research universities.
uOttawa focuses research strengths and efforts in four Strategic Areas of Development in Research (SADRs):
With cutting-edge research, our graduate students, researchers and educators strongly influence national and international priorities.
The Faculty of Science has become a true centre of excellence in research through its world-class professors as well as its programs and infrastructure in Biology, Chemistry, Earth Sciences, Mathematics and Statistics, and Physics.
The research accomplished by its 140 internationally recognized professors, its approximately 400 graduate students and its dozens of postdoctoral researchers and visiting scientists has positioned the Faculty of Science as one of the most research intensive science faculties in Canada. Our professors have received many international and national awards including three NSERC Gerhard Herzberg Gold Medal winners and numerous Fellows of the Royal Society of Canada.The Faculty of Science, through its strategic use of infrastructure programs, hosts world-class Core Facilities and is at the leading edge for the study of Catalysis, Experimental and Computational Chemistry, Environmental Toxins, Nuclear Magnetic Resonance, Isotope Analysis, Molecular Biology and Genomics, X-Ray Spectrometry/Diffractometry, Geochemistry, Mass Spectrometry, Physiology and Genetics of Aquatic Organisms, and Photonics. The Faculty is also associated with the Fields Institute for research in mathematical science and the Centre de recherche mathématiques (CRM) at the Université de Montréal, providing a unique setting for mathematical research.
For more information, refer to the list of faculty members and their research fields on Uniweb .
IMPORTANT: Candidates and students looking for professors to supervise their thesis or research project can also consult the website of the faculty or department of their program of choice. Uniweb does not list all professors authorized to supervise research projects at the University of Ottawa.
Not all of the listed courses are given each year. The course is offered in the language in which it is described.
A 3-unit course at the University of Ottawa is equivalent to a 0.5-unit course at Carleton University.
BIO 5101 Topics in Biotechnology (3 units)
A course concerned with the utilization of biological substances and activities of cells, genes and enzymes in manufacturing, agricultural and service industries. A different topic will be selected each year. This course is equivalent to BIOL 5001 at Carleton University.
Course Component: Lecture
Prerequisite: A course in cell physiology or biochemistry, or permission of instructor.
BIO 5102 Advanced Field Ecology (3 units)
Field experience in a new environment (e.g. local, national, international) to learn about ecological processes (note extra fees associated with course). This course is equivalent to BIOL 5605 at Carleton University.
BIO 5104 Advances in Applied Biochemistry (3 units)
Contemporary methods of recombinant DNA technology combined with modern methods and strategies for expressing, secreting, purifying and characterizing proteins. This course is equivalent to BIOL 5004 at Carleton University.
BIO 5105 Advanced Neuroethology (3 units)
A comparative and evolutionary approach to studying neural mechanisms underlying animal behaviour, including genetic, neural and hormonal influences on behaviour. This course is equivalent to BIOL 5801 at Carleton University.
Prerequisites: Biology 61.335 and 61.361 or equivalents and registration in a graduate program, or written permission of the department.
BIO 5107 Seminar in Biochemistry I (3 units)
A graduate seminar on current topics in the field of Biochemistry. This course introduces the seminar format and involves student, faculty and invited seminar speakers. The student will present a seminar and submit a report on a current topic in Biochemistry. This course is equivalent to BIOL 5002 at Carleton University. Includes: Experiential Learning Activity
BIO 5108 Bayesian Statistics for Biologists (3 units)
Introduction to the philosophy of Bayesian inference; practical experience applying to biological data. Model formulation, identification of appropriate priors and resulting posteriors given priors and data, and the practice of drawing inferences from these posteriors. This course is equivalent to BIOL 5408 at Carleton University.
BIO 5109 Biological Data Science in R (3 units)
Develops the practical skills needed to work with large and complex datasets, as a complement to statistical methods. Topics include programming, quality control, tidy data, visualization, project organization, reproducibility, how to troubleshoot code, and how to translate research goals into a project pipeline. This course is equivalent to BIOL 5404 at Carleton University.
BIO 5111 Biophysical Techniques (3 units)
Theory and application of current biochemical/biophysical instrumentation and techniques including X-ray crystallography, nuclear magnetic resonance spectrometry, infrared, circular dichroism and fluorescence spectroscopy, isothermal titration and differential scanning calorimetry. This course is equivalent to BIOL 5111 at Carleton University.
BIO 5126 Analysis of Next-Generation Sequence Data (3 units)
Assembly and analysis of next-generation sequence (NGS) data. Through hands-on exercises and independent projects, students will learn to use tools for quality control, assembly, mutation calling, and other NGS applications. No previous knowledge of bioinformatics or programming is required. This course is equivalent to BIOL 5526 at Carleton University.
BIO 5128 Molecular Methods (3 units)
An intensive two-week laboratory course where students are introduced to methods such as CRISPR-Cas9 genome editing, in situ hybridization, immunohistochemistry, qRT-PCR and digital droplet PCR.
Course Component: Theory and Laboratory
BIO 5130 Ethnobotany and Ethnopharmacology (3 units)
Introduction and current perspectives on world ethnobotanies, traditional knowledge, medicinal and food systems; quantitative and qualitative methods; ethical requirements; pharmacological basis of traditional drugs, phytochemsitry, drug discovery and development; safety, risk assessment and regulations.
BIO 5144 Plant Molecular Biology (3 units)
An introduction to plant gene structure and function, cloning into plants and the manipulation of plant genes. The course will combine elements of plant biochemistry, physiology and molecular biology with a strong emphasis on practical research aspects. This course is equivalent to BIOL 5144 at Carleton University.
BIO 5158 Applied Biostatistics (3 units)
Applied biostatistics to real problems. Experimental design and data collection. Consequences of violating assumptions of different tests. Monte Carlo and Bootstrap analysis. Case studies and exercises in using statistical analysis packages. This course is equivalent to BIOL 5158 at Carleton University.
BIO 5302 Methods in Molecular Genetics (3 units)
Theory and associated applications of emerging methods in molecular genetics, including information gathered from large-scale genome-wide analysis and protein-protein interaction data, and how this information can advance understanding of cell biology. This course is equivalent to BIOL 5105 at Carleton University.
Prerequisites: Graduate standing and permission of the department.
BIO 5305 Biostatistics I (3 units)
Application of statistical analyses to biological data. Topics include ANOVA, regression, GLMs, and may include loglinear models, logistic regression, general additive models, mixed models, bootstrap and permutation tests. This course is equivalent to BIOL 5407 at Carleton University.
Prerequisites: Graduate standing, courses in elementary ecology and statistics and permission of the department.
BIO 5306 Modelling for Biologists (3 units)
Use and limitations of mathematical and simulation modelling approaches for the study of biological phenomena. This course is equivalent to BIOL 5409 at Carleton University.
BIO 5308 Laboratory Techniques in Molecular Genetics (3 units)
Laboratory course designed to give students practical experience in recent important techniques in molecular genetics. This course is equivalent to BIOL 5106 at Carleton University.
BIO 5310 Advanced Evolutionary Biology (3 units)
Advances in micro-and macroevolution including the mechanisms both driving and constraining evolutionary change, phylogenetic relationships, patterns of evolutionary change at the molecular or phenotypic level, and evolutionary theory and techniques as applied to these areas. This course is equivalent to BIOL 5510 at Carleton University.
BIO 5311 Advanced Evolutionary Ecology (3 units)
The ecological causes and consequences of evolutionary change, focussing on how the ecological interactions among organisms and their biotic and abiotic environments shape the evolution of phenotypic and species diversity. This course is equivalent to BIOL 5511 at Carleton University.
BIO 5312 Principles and Methods of Biological Systematics (3 units)
Biological systematics with reference to morphological and molecular character evolution and phylogeny reconstruction.
BIO 5313 Topics in Evolutionary and Comparative Biology (3 units)
In the ever-diversifying field of evolutionary and comparative biology, it is becoming increasingly necessary for early-career researchers to cultivate a broad set of skills, eventually to launch a project of interdisciplinary nature. This course provides workshop and hands-on training for students to develop broad basis and familiarity with the research toolkit of modern biology. Topics include the use of statistical programs, 3D data acquisition and analysis, cladistic analysis and phylogenetic comparative method, microscopy and histology, basic bioinformatics, and scientific illustration. Each workshop will be led by a faculty expert. Offered in alternate years. This course is equivalent to BIOL 5313 at Carleton University.
BIO 5314 Advances in Aquatic Sciences (3 units)
Advanced theoretical and applied aquatic sciences including current topics in limnology and oceanography (e.g. impacts of climate change, invasive species, and atmospheric pollution) with implications for lake, river, coastal and wetland management. This course is equivalent to BIOL 5514 at Carleton University.
BIO 5320 Advances in Conservation Biology (3 units)
Interdisciplinary exploration of the science of scarcity and diversity in a human dominated world. This course is equivalent to BIOL 5520 at Carleton University.
BIO 5558 Biostatistiques appliquées (3 crédits)
Application des biostatistiques à des problèmes concrets. Design expérimental et échantillonnage. Impact des violations des hypothèses implicites d'application de divers tests. Analyse de Monte Carlo et Bootstrap. Études de cas et exercices d'utilisation de logiciels courants d'analyse statistique.
Volet : Cours magistral
BIO 5950 Recherche et communication en biologie / Research and Communication in Biology (3 crédits / 3 units)
Une introduction aux études supérieures en biologie en mettant l'accent sur les compétences centrales requises pour mener à bien des projets de recherche et développer les habiletés professionnelles requises. Le cours s'adresse aux étudiant.e.s débutant.e.s aux cycles supérieurs et sera composé de modules couvrant la rédaction scientifique et la communication orale, la gestion de projets de recherche, le développement de carrière et diverses compétences centrales requises dans les programmes d'études supérieures en biologie. / An introduction to graduate studies in biology with an emphasis on central skills required to successfully conduct research projects and develop required professional abilities. The course is intended for starting graduate students and will be composed of modules covering scientific writing and oral communication, research project management, career development and various central skills required in biology graduate programs.
Volet / Course Component: Séminaire / Seminar
BIO 6103 Special Topics in Neuroscience (3 units)
An in-depth study of current topics in neuroscience. Course content varies yearly and has recently included cognitive neuroscience, neuropharmacology, neurodegeneration, and behavioural medicine. Also listed as PSYC 6300. This course is equivalent to BIOL 6203 at Carleton University.
BIO 6300 Advanced Science Communication (3 units)
The theory and practice of effective science communication. Topics may include : writing for, presenting to, and engaging with diverse audiences, as well as graphic design and data visualization, social and digital media, and knowledge mobilization. Experiential Learning Activity: Applied Research. This course is equivalent to BIOL 6500 at Carleton University.
BIO 6304 Techniques in Neuroscience (3 units)
Completion of a research project carried out under the supervision of a neuroscience faculty member. The student will learn a new neuroscience technique and apply it to a research objective. May be repeated for different projects. Also listed as PSYC 6204. This course is equivalent to BIOL 6204 at Carleton University.
BIO 6305 Advanced Seminar in Neuroscience (3 units)
A comprehensive pro-seminar series, covering issues ranging from cellular and molecular processes through to neural systems and behaviours as well as psychopathology. Also listed as PSYC 6202. Courses BIO 6305 , BIO 6303 (BIOL 6303) cannot be combined for units. This course is equivalent to BIOL 6305 at Carleton University.
BIO 8102 Special Topics in Biology (3 units)
Selected aspects of specialized biological subjects not covered by other graduate courses. This course is equivalent to BIOL 5502 at Carleton University.
Course Component: Laboratory, Lecture
BIO 8105 Advances in Applied Ecology (3 units)
The application of ecological and evolutionary principles in addressing resource management challenges and environmental problems. This course is equivalent to BIOL 5512 at Carleton University.
Permission of the Department is required.
BIO 8108 Advanced Topics in Development (3 units)
Recent advances in developmental biology. Topics may include embryonic induction, regulation of morphogenesis and differentiation, mechanisms of regional specification and pattern formation, and developmental genetics. This course is equivalent to BIOL 6505 at Carleton University.
BIO 8109 Advanced Molecular Biology (3 units)
In-depth coverage of the structure, function, and synthesis of DNA, RNA, and proteins. This course is equivalent to BIOL 6001 at Carleton University.
BIO 8115 Genomics in Graduate Studies (3 units)
Applying tools of genomics in the current research environment. Students will build an original research proposal that includes genomics analyses distinct from those they currently use. The goal of this course is to investigate how genomics (broadly defined) can help students tackle and/or uncover new interesting questions related to their current graduate research. This course is equivalent to BIOL 6115 at Carleton University.
BIO 8116 Advances on Plant Molecular Biology (3 units)
Use of molecular genetics in general plant biology and the contribution of plant genomics to our understanding of plant metabolism, plant development, and plant interactions with the environment at the molecular, genome, and cellular levels. This course is equivalent to BIOL 6002 at Carleton University.
Prerequisite: BIO 8109 /61.601F1 and this course normally will be offered together in the same year but only in alternate years.
BIO 8120 Directed Studies in Biology (3 units)
One-on-one instruction in selected aspects of specialized biological subjects not covered by other graduate courses. Students may not take this course from their thesis supervisor(s), and are limited to one directed studies course per program. This course is equivalent to BIOL 5502 at Carleton University.
BIO 8122 Advanced Insect Biology (3 units)
Overview of the biological processes that allow insects to function in their environments and to overcome the constraints and limitations that the environment places on them. This course is equivalent to BIOL 5307 at Carleton University.
Prerequisite: In addition to the course material, students will write two terms papers (Alter nate years).
BIO 8162 Advanced Endocrinology (3 units)
Major topics in comparative endocrinology: understanding the structure, function and evolution of vertebrate endocrine systems, including endocrine disruption. This course is equivalent to BIOL 5402 at Carleton University.
Prerequisite: An undergraduate Endocrinology course ( BIO 4127 or equivalent).
BIO 8303 Advanced Microscopy (3 units)
Development of the practical skills of microscopy through original research and supporting theory lectures. This course is equivalent to BIOL 5203 at Carleton University.
Prerequisites: Open to 4th year and graduate students with consent of the instructor.
BIO 8320 Advanced Plant Biology (3 units)
Recent developments in plant biology. Topics may include plant anatomy, systematics, evolution, genetics, ecology, ethnobotany, cell biology, and/or biotechnology. This course is equivalent to BIOL 6300 at Carleton University.
Prerequisite: Biology 61.425 and Biology 61.426/427, or permission of the department.
BIO 8361 Advanced Animal Physiology (3 units)
Recent advances in animal physiology, emphasizing comparative, evolutionary and environmental approaches. This course is equivalent to BIOL 6304 at Carleton University.
BIO 8365 Advanced Behavioural Ecology (3 units)
Recent advances in behavioural ecology including topics such as the evolution of tactics and strategies of group living, foraging, anti-predation, resource use and defence, cooperation, reproduction, and parental care. This course is equivalent to BIOL 5802 at Carleton University.
BIO 8510 Thèmes choisis en biologie (3 crédits)
Aspects de sujets biologiques spécialisés qui ne sont pas couverts dans d'autres cours d'études supérieures.
BIO 8520 Études dirigées en biologie (3 crédits)
Enseignement individualisé sur un sujet biologique spécialisé qui n'est pas couvert dans d'autres cours d'études supérieures. Il est interdit de suivre ce cours avec son directeur de thèse. Limite d'une seule étude dirigée par programme.
BIO 8910 Thèmes choisis en biologie / Special Topics in Biology (3 crédits / 3 units)
Aspects de sujets biologiques spécialisés qui ne sont pas couverts dans d'autres cours d'études supérieures. / Selected aspects of specialized biological subjects not covered by other graduate courses.
Volet / Course Component: Cours magistral / Lecture
Prérequis : connaissance passive de l'anglais. / Prerequisite: Passive knowledge of French.
BIO 8938 Interaction entre plantes et animaux / Plant Animal Interactions (3 crédits / 3 units)
Les substances métaboliques secondaires des plantes et leur rôle en tant que phagorépresseurs ou phagostimulants pour les animaux et en tant qu'agents antifongiques ou allélopathiques. On discutera de la co-évolution des plantes et des organismes phytophages (insectes et mammifères) et des dimensions physiologique et écologique de cette relation. / Secondary metabolites of plants and their role as attractants or antifeedants to animals and as allelopathic or antifungal agents. Emphasis will be placed on co-evolution of plants and phytophagous organisms such as insects and mammals, and the ecological and physiological dimensions of this relationship. Offered in alternate years. Ce cours est équivalent à BIOL 6404 à la Carleton University. / This course is equivalent to BIOL 6404 at Carleton University.
BIO 8940 Statistiques avancées et science ouverte / Advanced Statistics and Open Science (3 crédits / 3 units)
Les analyses statistiques sont fondamentales à un processus scientifique rigoureux. Par conséquent, il est primordiale de comprendre les statistiques et de reporter correctement les analyses pour améliorer la transparence et la qualité de la science. Le cours a pour objectifs: 1) d'améliorer la compréhension des modèles statistique avancés (incluant les modèles mixtes généralisés); 2) de développer de bonnes habitudes pour coder (utilisation de R et Rmarkdown); 3) d'améliorer la gestion des données et du code statistique (manipulation de données et github); et 4) de présenter les principes de science ouverte (se basant sur OSF). / Statistics are a key component of rigorous science and as such there is a need to both understand advanced statistics and properly document the analysis to improve scientific communication transparency and quality. The course aims to 1) provide an understanding of advanced statistical models (including generalized linear mixed models), 2) develop good coding practices (using R and Rmarkdown), 3) improve data and code management (data manipulation and github) and 4) present the principles of open science (using OSF).
BIO 8950 Recherche et communication en biologie / Research and Communication in Biology (3 crédits / 3 units)
BIO 9998 Examen de synthèse / Comprehensive Examination
Volet / Course Component: Recherche / Research
For more information about undergraduate studies at the University of Ottawa, please refer to your faculty .
For more information about graduate studies at the University of Ottawa, please refer to your academic unit .
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We are looking for two PhD students to investigate the effects of irradiation on cell metabolism and reciprocally.
1. Characterization of tumour cells exposure to radiations and combined influence of environmental acidity PhD Project TIMC-IJCLab https://mycore.core-cloud.net/index.php/s/XILL8bZGrmWPOws Application deadline : 30/06/2024
2. Mathematical and computational approaches to explore the reciprocal interaction between cell metabolism and radiation therapy in cancer PhD Project TIMC-Swansea https://mycore.core-cloud.net/index.php/s/L0KEkPy0wfN0Uzs Application deadline : 05/06/2024
Both PhD Projects are fully funded and are due to start on October 1st, 2024.
A 18-months Postdoc position is available in the Laboratoire de Mathématiques de Besançon (LMB), at the University of Bourgogne Franche-Comté, France, to work on the project “Parameter estimation for nonlocal PDEs with applications in cell biology”.
We are looking for candidates with a strong research background in either one of the 2 research directions of this project: (a) parameter identification and uncertainty quantification for time-evolving PDEs, and/or (b) numerical analysis (a posteriori error estimates and adaptive mesh refinement) for time-evolving PDEs.
Applications will be accepted on a rolling basis, and their review will continue until the position is filled. The contract will start as soon as possible.
For more information, please contact Raluca Eftimie ( [email protected] ). Applicants are requested to send their expression of interest electronically in a single PDF file. Applications should include (1) a cover letter, (2) a CV, (3) contact details of up to three referees.
I am looking for motivated master interns, PhD students, and postdocs in theoretical physics or applied mathematics to work on the interactions between mechanics, biochemistry, and genetics in stem cell aggregates.
The positions are fully funded by the ERC Synergy Grant project BREAKDANCE. Our theory work will be carried out in close collaboration with the experimental labs of Pierre-François Lenne (Marseille), Vikas Trivedi (Barcelona), and Verena Rupprecht (Barcelona).
For more details on the positions, please see https://www.merkellab.net/open-positions
A recent preprint from our consortium can be found here: https://www.biorxiv.org/content/10.1101/2023.09.22.559003
The start date is flexible.
Please do not hesitate to contact me for any informal inquiries: [email protected]
The Department of Computational and Quantitative Biology (LCQB) at the prestigious Sorbonne University, located in the heart of Paris, France, is on the lookout for a new director set to start January 1, 2025. This position is for a five- year term, with the possibility of renewal.
Full details are available here: https://dropsu.sorbonne-universite.fr/s/iegn9ftgTb6EmbS
4-years position (75%) | E13 TV-L | Potsdam close to Berlin
Project Title: Modelling context-dependent shifts in the forms of symbioses
Project Description: Symbiotic interactions - representing prolonged physical associations of several species - are common in natural systems and can determine population dynamics, species persistence and ecosystem functioning, as demonstrated for example in coral reefs and plantpollinator networks. Symbiotic interactions can take different forms including parasitism, mutualism and competition. Depending on the underlying costs and benefits of the symbiotic interaction, the form of symbiosis may shift between different types of species interactions, e.g. mutualism and predator-prey interaction (see figure). While recent work has shown that the costs and benefits of symbioses depend on the densities of the symbiosis partners, we currently lack an understanding of how the form of symbioses depends on species traits and the overall food web context. This is particularly important as individuals and populations may adapt their traits to altered environmental conditions and as the food web structure may strongly vary across time and space. Hence, we want to improve general theory in community ecology by accounting for context-dependent changes in the form of symbiosis including the species’ potential to adapt to altering conditions in a food web context.
The project is embedded into a network of experimental and theoretical working groups across Germany, conducting regular workshops and meetings promoting international networking. The prospective PhD student will join the working group of Prof. Dr. Ursula Gaedke, jointly supervised by Dr. Toni Klauschies. The prospective PhD candidate has the opportunity to closely interact with an experimental PhD student addressing the same overarching topic in our working group and related research groups on Campus. They can join the Potsdam Graduate School (PoGS) allowing for a broad interdisciplinary training of soft skills and early career development. The unique location of our campus in Park Sanssouci is part of a historical place in Potsdam providing a fruitful scientific environment and good working atmosphere.
Your qualification: MSc degree in Ecology or other relevant subjects such as Physics, Mathematics or Environmental Science. We are seeking enthusiastic and committed candidates who enjoy ecology and applied mathematics with a solid background in ecology and ecological modelling. The successful candidate is expected to implement and analyze numerically differential equation models with a modern programming language such as Python, MatLab or Mathematica. Very good English writing and communication skills are expected, German is an asset but not essential.
Application: To apply, please send the following documents as a single PDF to [email protected] (the position is open until it is filled)
· Cover letter, including a statement of motivation and from when on you would be available · Detailed curriculum vitae including a description of your pre-knowledge in (theoretical) ecology, programming and former research activities · Certifications of education · If possible, provide letters of recommendation from previous supervisors
4-years position (75%) | E13 TV-L | Potsdam close to Berlin
Project Title: Context dependent effects on protective symbiosis driven by predation, competition and parasitism
Project Description: Symbiosis is a biological interaction which refers to the physical association of individuals from different species that can be positive, negative or neutral to either species. In planktonic systems several symbiotic interactions can occur simultaneously and can interact with each other. We want to study the symbiosis between Daphnia and the rotifer Brachionus rubens within a multi-species interaction web. B. rubens can attach to Daphnia which reduces the fitness of Daphnia. At the same time both species compete for resources (algae). The attachment becomes more relevant, when a predator of B. rubens, A. brighwelli (another rotifer) is present because attached B. rubens are protected from predation. The aim is to quantify the costs and benefits for Brachionus and Daphnia under varying conditions and to better understand symbioses in a community context. Thus, the present project will contribute to a new food web theory which includes density dependence of symbioses. In addition, video analyses will be performed to study the attachment process in more detail.
In the case of final approval, the project is embedded into a network of several experimental and theoretical working groups across Germany conducting regular workshops and meetings.
The prospective PhD student will join the working group of apl. Prof. Dr. Guntram Weithoff at the University of Potsdam. We provide an active research environment covering a broad range in ecological research. Beyond that, the Potsdam Graduate School (PoGS) offers a broad program on interdisciplinary training in soft skills and early career development.
Your qualification: MSc degree in Ecology preferably in Aquatic Ecology or a related field. We are looking for a motivated candidate with good experimental skills. Further relevant skills are: sound knowledge in conceptional ecology, statistics and very good English in writing and communication.
Application: To apply, please send the following documents as a single PDF to [email protected] (the position is open until it is filled): • Cover letter, including a statement of motivation and from when on you would be available • Curriculum vitae including information on relevant skills • Certificates of education • If possible, provide letters of recommendation from previous supervisors
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Are you a high-achieving student interested in pursuing a PhD in molecular and biomedical sciences? Do you want to work with top researchers, in a world-class research facility, in a beautiful city at the bottom of the world? Then check out the Division of Biomedical Science and Biochemistry at the Australian National University.
Our Science : We provide an outstanding research environment with a world-class reputation. We are an interdisciplinary division, with highly integrated labs that work at the forefront of membrane transport processes, host-pathogen biology, drug and vaccine development, immune processes, cancer biology and evolutionary genomics. We research a wide range of topics – from the physiology of intracellular parasites to the assembly of bacterial nanomachines; from the structure and biophysics of membrane transporters to the early evolution of animals. If you are interested in doing a PhD with us check out our Research Groups below, and contact prospective supervisors.
Our Values : Our division includes award-winning supervisors and teachers, and we pride ourselves on providing high-quality supervision to our students. Our PhD students are well supported through internal funding, including for conference travel and computational resources, and our labs are located in a well-equipped, modern research facility. We have a thriving community of PhD students, and our graduates go on to productive careers in many areas of science and beyond. We value diversity and inclusivity in science and have active policies to prevent discrimination. Our faculty and their research teams are comprised of people from all over the globe and from all walks of life.
Location : The ANU is situated in Canberra, Australia’s capital city. Canberra is a well-resourced regional city, set amongst beautiful mountains and surrounded by eucalypt forests. Bike riding and hiking are everyday activities, and we are a two-hour drive away from both winter snowfields and some of the most beautiful beaches on Earth. Canberra’s birdlife is stunning, and we regularly have to brake for kangaroos on our daily commute. Despite its regional setting, Canberra is a vibrant, multicultural city. There are frequent cultural evenings, festivals, art exhibits, music events, and world-class restaurant and coffee scenes.
The Australian National University: We are a research-intensive university and have an international reputation for research excellence. The ANU is frequently ranked as Australia’s top university, and one of the top 50 universities in the world.
Candidate eligibility : Domestic applicants must be Australian citizens or permanent residents, or New Zealand citizens, and must have previously completed a 1 st class Honours or Masters thesis in a relevant field. Fully funded International PhD scholarships are highly competitive at the ANU. To be a viable candidate you need: (1) excellent undergraduate marks; (2) a Masters by Research with a high mark; or (rarely) equivalent proof of research experience. (3) While not essential, having published research papers in the past is an advantage.
What do I do? First, contact potential supervisors from the list of staff on the Research Groups above (include your CV and grades, and some details about what it is that attracts you to their field of research). If they are interested in supervising you, you can submit an application to commence a PhD via the online link below. Your application should include a detailed CV, full academic transcripts, the name of your potential supervisor, and a brief description of a research project that you and your potential supervisor have discussed. Entry to the PhD program is open to applicants with a Bachelor degree that have also completed (or are in the process of completing) an Honours or Masters research degree that comprises at least a half year, full time research component and a thesis (5,000-10,000 words). Applicants with significant research experience and publications may be deemed eligible if their achievements can be justified as equivalent to completing an Honours/Masters degree. As part of your application for undertaking a PhD, you will automatically be considered for a stipend scholarship if you tick the scholarship box in the application. Your potential supervisor or the Biomedical Science and Biochemistry graduate convenor, Dr Giel van Dooren, can advise you on how competitive your application is likely to be for obtaining a stipend scholarship. If you have questions about any aspect of the PhD application process, you can contact Giel van Dooren ( [email protected] )
The application form is here , along with general information on how to apply and the details about the Doctor of Philosophy program . PhD applications are due by 15 April (the mid-year round for both international and domestic students), 31 August (international students), or 31 October (domestic students). Your application will require:
* If you are an international student and have the possibility of PhD funding from your home country or elsewhere, let a potential supervisor know.
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ESMTB annually honours the best PhD thesis in the field of mathematical and theoretical biology with the Reinhart Heinrich Doctoral Thesis Award, and together with the SFBT awards the Ovide Arino Outreach Award every two years for seminal PhD contributions in mathematical and theoretical biology from southern countries. ESMTB is a nonprofit ...
The Systems, Synthetic, and Quantitative Biology PhD Program aims to explain how higher level properties of complex biological systems arise from the interactions among their parts. This field requires a fusion of concepts from many disciplines, including biology, computer science, applied mathematics, physics and engineering.
The Journal of Theoretical Biology is the leading forum for theoretical perspectives that give insight into biological processes. It covers a very wide range of topics and is of interest to biologists in many areas of research, including: • Brain and Neuroscience. • Cancer Growth and Treatment.
Biology, PhD. The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student's thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology ...
The Department of Theoretical Biology uses mathematics and computer simulations to study the dynamics of evolution. Our team is interdisciplinary, with backgrounds ranging from biology, physics, and mathematics to computer science and economics. ... If you want to work with Arne Traulsen as a PhD candidate or PostDoc, please inquire directly ...
The evolutionary biology PhD is a transdisciplinary graduate degree program that provides doctorate-level training in the historical, conceptual, empirical and quantitative aspects of biological evolution. Evolution is a fundamental scientific concept underlying all aspects of modern biological, environmental and health-related research.
Professor of Cell Biology. View Full Profile. Biological and Biomedical Sciences Program (BBS) Yale University. New Haven, CT 06520-8084. you could attend in June 2024. Development of novel theoretical principles of biology, predictive mathematical modeling of biological processes, direct comparison between quantitative models.
The Biophysics Graduate Program offers seven areas of research emphasis: 1. Biophysical approaches to cell biology. The fields of molecular and cellular biology are exceptionally strong at UCSF, and many faculty members in these areas use cutting-edge tools of biophysics. These approaches include quantitative analyses of chaperones, protein ...
The Department of Theoretical Biology uses mathematics and computer simulations to study the dynamics of evolution. Our team is interdisciplinary, with backgrounds ranging from biology, physics, and mathematics to computer science and economics. Our goal is, on the one hand, to model biological systems with a close connection to empirical ...
Theoretical biology is trying to do just that; and the fact that in doing so many concepts of the physical sciences are "taken over" and used to describe biological phenomena in an abstract manner provides an argument for considering theoretical biology as a legitimate domain in biophysics. Its aim, of course, is the unification of ...
Computational and Theoretical Biology. Prof. Jan Pieter Abrahams. Uncovering the fundamental shapes of life. Prof. Attila Becskei. Mechanisms controlling gene regulatory networks. Prof. Flavio Donato. Development of neuronal circuits and cognitive functions. Prof. Knut Drescher. Development and functions of bacterial communities.
HMS New Research Building, Room 466D (4th Floor) 77 Avenue Louis Pasteur, Boston, MA 02115. [email protected]. p: 617 525-4725. George M. Church, Ph.D. (He/ Him/ His) Robert Winthrop Professor of Genetics, & Health Sciences & Technology, Harvard Medical School/ MIT.
Theoretical Biophysics. Biophysics is an interdisciplinary science that applies the approaches and methods of physics to study biological systems. Biophysics covers all scales of biological organization, from molecular to organismic and populations. To most physicists, almost everything about evolution is puzzling.
The track Theoretical Biology and Bioinformatics in the master program Biology and Biocomplexity of the Graduate School of Life Sciences provides an optimal preparation for a PhD in Computational Life Sciences. Students with a master education in mathematics or physics can also be admitted by taking additional courses in the relevant biological ...
David PENNY, Professor of theoretical biology | Cited by 20,024 | of Massey University, Palmerston North | Read 337 publications | Contact David PENNY
The Center offers many research opportunities to postdoctoral fellows and graduate and undergraduate students. These Opportunities for Research in Biophysics, Informatics and Theoretical Science (ORBITS), encourage using both theoretical and experimental physics to further the understanding of biology and its application to biomedicine.
PhD position in Theoretical Biology. When. 31 Jul 2022. Location. University of Münster, Germany. I am looking for a PhD student to work in theoretical evolutionary genetics. The PhD project will explore potential conflict and collaboration between transposons and their Drosophila hosts. To this end the PhD student will model population ...
Degree offered: Doctorate in Philosophy (PhD) Registration status option: Full-time. Language of instruction: English. Program option (expected duration of the program): with thesis (12 full-time terms; 48 consecutive months) Academic units: Faculty of Science, Department of Biology, Ottawa-Carleton Institute of Biology. Program Description.
The graduate program in Structural Biology is intended to prepare students for careers as independent investigators in cell and molecular biology. The principal requirement of a PhD degree is the completion of research constituting an original and significant contribution to advancing knowledge. It is training in a major with connections to ...
PhD position in the Theoretical Biology group at Bielefeld University: 29 Apr 2022 PhD scholarship: Mathematical modeling in infection and immunity: 25 Apr 2022 Two 3-Year Post-Doc positions at the interface of developmental biology and mathematical modelling: 20 Apr 2022 PhD project in Systems Biology
Learn more about Theoretical Chemistry and Biology (PhD) program with KTH Royal Institute of Technology including the program fees, scholarships, scores and further course information
Stuart Alan Kauffman (born September 28, 1939) is an American medical doctor, theoretical biologist, and complex systems researcher who studies the origin of life on Earth.He was a professor at the University of Chicago, University of Pennsylvania, and University of Calgary.He is currently emeritus professor of biochemistry at the University of Pennsylvania and affiliate faculty at the ...
The "Theoretical Biology" series publishes volumes on all aspects of life sciences research for which a mathematical or computational approach can offer the appropriate methods to deepen our knowledge and insight. Topics covered include: cell and molecular biology, genetics, developmental biology, evolutionary biology, behavior sciences ...
Our PhD students are well supported through internal funding, including for conference travel and computational resources, and our labs are located in a well-equipped, modern research facility. We have a thriving community of PhD students, and our graduates go on to productive careers in many areas of science and beyond.
229 Gilmer Hall University of Virginia P.O. Box 400328 Charlottesville, VA 22904-4328 434-982-5474 www.bio.virginia.edu
Cancer Biology; Cell Adhesion and Cytoskeleton; Cellular and Molecular Metabolism; Developmental Biology and Regenerative Medicine; Immunology; Membrane Biology and Protein Trafficking; Molecular and Genome Biology of Microbes; Physiology; Plant Biology; RNA Biology; Systems Biology; Transcriptional Mechanisms; Virology; Prospective Students
The interactions between chromatin and the nuclear lamina orchestrate cell type-specific gene activity by forming lamina-associated domains (LADs) which preserve cellular characteristics through gene repression. However, unlike the interactions between chromatin segments, the strength of chromatin-lamina interactions and their dependence on cellular environment are not well understood. Here ...
Cancer Biology; Cell Adhesion and Cytoskeleton; Cellular and Molecular Metabolism; Developmental Biology and Regenerative Medicine; Immunology; Membrane Biology and Protein Trafficking; Molecular and Genome Biology of Microbes; Physiology; Plant Biology; RNA Biology; Systems Biology; Transcriptional Mechanisms; Virology; Prospective Students
Michelle Maillet (Academic Associate), Lauren Kay (Clerical), Chantal Marotte (Management), and Moshe Dalva (Technical) have been named recipients of the Faculty of Science Excellence Awards. These awards, announced at the Faculty of Science Council meeting on May 21st, recognize the outstanding contributions made by members of the Faculty's administrative and support staff during the 2023 ...
Cancer Biology; Cell Adhesion and Cytoskeleton; Cellular and Molecular Metabolism; Developmental Biology and Regenerative Medicine; Immunology; Membrane Biology and Protein Trafficking; Molecular and Genome Biology of Microbes; Physiology; Plant Biology; RNA Biology; Systems Biology; Transcriptional Mechanisms; Virology; Prospective Students