phd biochemistry stanford

• Office of Graduate Education

Program Overview

When you join Stanford Biosciences, you join a collaborative network tackling some of the world’s toughest questions.  The Stanford Biosciences  Home Programs  comprise nine departments and five interdisciplinary programs, which span the School of Medicine and the School of Humanities and Sciences.  These Home Programs are the foundation of our collaborative culture, offering students the opportunity to tailor their graduate education  by working within an entire network of faculty, labs, and approaches to pursue their research.

Each student is admitted to a particular Home Program and initiates training with a core group of faculty, students, and postdoctoral fellows who share scientific interests. Many Home Programs host annual retreats—facilitating the exchange of ideas between Stanford colleagues and fostering team-building—as well as seminar series that invite outside speakers.

In addition to that intimate setting, all Biosciences students have access to faculty in every Home Program for laboratory rotations and potential thesis work.  One of Stanford Biosciences’ biggest strengths is the physical proximity of programs and labs , encouraging face-to-face collaboration and feeding an environment of interdisciplinary innovation. Indeed, the Biosciences PhD Programs combine the supportive atmosphere of a small program with the many opportunities afforded by a large umbrella program—the best of both worlds.

A closer look

The 14 Home Programs in Stanford Biosciences’ collaborative network:

Biochemistry

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Biomedical Data Science

Cancer Biology

Chemical and Systems Biology

Developmental Biology

Microbiology and Immunology

Molecular and Cellular Physiology

Neurosciences

Stem Cell Biology and Regenerative Medicine

Structural Biology

Related programs

Bioengineering.

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Biomedical Physics

Health Policy

Epidemiology and Clinical Research

Dual-Degree Programs

Providing a select group of medical students with an opportunity to pursue a training program designed to equip them for careers in academic investigative medicine.

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Ph.D. Program

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• Ph.D. Courses
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Advances In Basic Science And Molecular Medicine

The Department of Chemical and Systems Biology explores the frontiers of basic science and molecular medicine, particularly at the crossroads of cellular, chemical, and computational biology. We train Ph.D. students to apply genetic, chemical, cell biological, and quantitative methods to decipher the complex regulatory systems associated with normal physiology and disease states.

Specific research areas include cell signaling pathways, cell cycle control, epigenetics, cell fate specification, and genomic stability. The Chemical and Systems Biology Ph.D. program also emphasizes collaborative learning, and our research community includes scientists trained in molecular biology, cell biology, chemistry, physics, and engineering.

Our Ph.D. program consistently ranks among the top graduate training programs in the world. Most recently the National Research Council named us the top pharmacology-related training program in the United States, based on students’ GRE scores, faculty publications, median time to degree, program requirements, and training resources. The Chemical and Systems Biology graduate program was especially commended for the quality of its research activities.

Why Chemical And Systems Biology?

How do cells achieve directed migration? Why doesn’t a skin cell become a neuron? How do drug-resistant cancers arise and how might they be prevented or overcome? Finding answers to these and other biomedical questions increasingly requires molecular, quantitative, and interdisciplinary approaches.

The Department of Chemical and Systems Biology is uniquely focused on understanding cell biology at the molecular and systems levels, and many of its faculty have expertise in biochemistry, chemistry, physics, and engineering. Developing novel technologies for basic research and translating discoveries into therapeutic strategies are also areas of special interest in the Chemical and Systems Biology community.

Our goal is to train a new generation of scientists with the interdisciplinary skills and creative thinking required to tackle emerging challenges in biomedical research. We invite all interested students to apply to the Chemical and Systems Biology Ph.D. program through the Stanford Biosciences online application form. Applicants whose research interests match well with our scientific mission are encouraged to select Chemical and Systems Biology as their primary home program.

MCP-PHD - Molecular and Cellular Physiology (PhD)

Program overview.

The Department of Molecular and Cellular Physiology is in the Beckman Center for Molecular and Genetic Medicine.

A central goal of physiology in the post-genomic era is understanding how thousands of encoded proteins bring about the highly coordinated behavior of cells and tissues. Research in the department approaches this goal at many levels of organization, ranging from single molecules and individual cells to multicellular systems and the whole organism. The faculty share common interests in the molecular mechanisms of cell signaling and behavior, with a particular focus on structure/function analysis of ion channels and G-protein coupled receptors and their roles at the cellular, organ, and whole-organism levels; the molecular basis of sensory transduction, synaptic transmission, plasticity and memory; the role of ion channels and calcium in controlling gene expression in neural and immune cells; and the regulation of vesicle trafficking and targeting, cell polarity, and cell-cell interactions in the nervous system and epithelia. Research programs employ various approaches, including molecular and cell biology, biochemistry, genetics, biophysics, x-ray crystallography, solution NMR, electrophysiology, and in vitro and in vivo imaging with confocal and multi-photon microscopy.

The department offers required and elective courses for students in the School of Medicine and is also open to other qualified students with the consent of the instructor. Training of medical, graduate, and postdoctoral students is available. The program offers a course of study leading to a PhD degree. No BS is offered, and an MS is offered only in the unusual circumstance where a student completes the coursework, rotation, and the written section of the qualifying exam but cannot meet the requirements for the PhD.

Free Form Requisites

Candidates for PhD degrees at Stanford must satisfactorily complete a program of study that includes 135 units of graduate coursework and research.

Studying for the PhD is expected to occupy five years, including summers. The MCP course requirements for the program are as follows:

course Advanced Cell Biology 

course  MCP Bootcamp

course  MCP Journal Club and Professional Development Series

course  How Cells Work: Energetics, Compartments, and Coupling in Cell Biology

course  Foundations in Experimental Biology

BIOS 217 Foundations of statistics and reproducible research

course  The Responsible Conduct of Research, if funded on NSF or NIH training grants

·       Basic competency in molecular biology

This requirement can be met based on undergrad curriculum/research. If not completed, then the Stanford undergrad course BIO 83 can be taken to meet the requirement.

·      Selected courses 

Take the equivalent of two (2) courses from the following list, where mini-courses count as ½ course

o   BIOC 241 (Biological Macromolecules)

o   MCP 222 (Imaging: Biological Light Microscopy) or BIOPHYS 232 (Advanced Imaging Lab in Biophysics)

o   Minicourse BIOS 294 (Chemistry for Biologists and Others)

o   Minicourse BIOS 202 (Understanding Kinetics for Biologists and Biology)

o   Minicourse NEPR 204 (Neuroscience Molecular Core; requires instructor approval)

o   Minicourse NEPR 201 (Neuro-Cellular Core; requires instructor approval)

·       Advanced graduate courses or minicourses for a minimum of 6 units total. These courses do not need to be MCP courses but must be on a relevant scientific topic and approved by the Director of Graduate Studies.

Attend MCP Seminar Series

Attend the MCP Annual Retreat

Any additional courses required by your training grant or Qualifying Committee

Service Requirement: Minimum of 40 hours before 5th year of study (MCP Handbook)

Courses to meet program requirements must be taken for a letter grade, and students must earn a minimum grade of at least a B in every required course. Students must also maintain a minimum GPA of 3.0 by university policy. Failure to maintain the required grades and grade point average is evidence of unsatisfactory progress in the program.

Students should complete their required courses within the first two years of study. Exceptions may be made in cases where it was impossible to schedule courses because they were not offered within a student’s first two years. Students may petition the DGS for variances in the required courses, and such petitions may be granted in special circumstances where a student’s progress is otherwise exemplary. The DGS may consult with the MCP Graduate Program Committee on these variances.

Laboratory Rotations

In addition to the course requirements detailed above, a student is expected to complete laboratory rotations during the first year. While students typically explore three rotations, a student may opt to do a greater number of shorter rotations as long as the rotation process is completed by May 15 of the first academic year of study. The first two of these rotations must be in a laboratory within the department, while subsequent rotations may be in any laboratory within the Stanford Biosciences Program. The first rotation should last eight to ten weeks, and each subsequent rotation shall last no longer than eight weeks. Shorter rotations are encouraged as long as the total rotations are between 20 and 26 weeks. MCP requires a lab rotation evaluation to be completed by faculty member and student at the end of the rotation. Continuation of rotations beyond May 15 will require the approval of the Director of Graduate Studies. Additional rotations and/or time for rotations will be considered in special circumstances, but in all cases, students should plan to join a dissertation lab no later than the beginning of autumn quarter of the second year. Dissertation and University Oral Examination

The results of independent, original work by the students are presented in a dissertation. The oral examination is essentially a defense of the dissertation.

Qualifying Examination

All students in the program must pass a qualifying examination to advance to candidacy for the PhD. Students are expected to take the qualifying examination by the end of autumn quarter in the second year of study. Failure to take the qualifying exam by the end of autumn quarter of the second year of study is evidence of unsatisfactory progress in the program. If a student thinks they need additional time to schedule and take their exam, a request must be submitted in writing to the Director of Graduate Studies (DGS) by November 15 of autumn quarter. The DGS may grant additional time in compelling circumstances that do not indicate poor progress or may refer the matter to the graduate committee for further action.

Students are given two chances to pass the qualifying examination unconditionally. Failure to achieve an unconditional pass of the qualifying examination by the end of spring quarter of the second year is grounds for dismissal from the program.

Students form a qualifying examination committee consisting of at least three faculty members (members of the academic council, including the dissertation advisor), at least one of whom must be a member of MCP. This committee should be formed by the end of spring quarter of the first year of study. The Director of Graduate Studies should approve the composition of this committee. Students should also check with the department’s student services office to file all required paperwork by the end of spring quarter. The university maintains specific deadlines for filing for candidacy, and it is the student’s responsibility to be aware of them.

Advisors and Advisory Committees

A graduate advisory committee, currently professors Feng, Lewis, Maduke, and Madison, advises students during the period before the formation of their qualifying committees.

Financial Aid

Students may be funded by their advisors' research grants, training grants, department funds, or extramural funds. Students are encouraged to obtain funding from outside sources such as NIH and NSF.

Biochemistry

Christopher O. Barnes

Martha S. Cyert

Scott Dixon

Judith Frydman

Arthur Grossman

Ashby Morrison

Mary Beth Mudgett

Naima G. Sharaf

Jan Skotheim

Shannon Yan

ExploreDegrees Archive, 2011-12

Explore courses, alphabetical index.

Bulletin Archive

This archived information is dated to the 2011-12 academic year only and may no longer be current.

For currently applicable policies and information, see the current Stanford Bulletin .

Doctor of Philosophy in Biochemistry

Requirements for the M.S. and Ph.D. degrees are described in the " Graduate Degrees " section of this bulletin. The department does not offer undergraduate degrees.

The Department of Biochemistry offers a Ph.D. program which begins in the Autumn Quarter of each year. The program of study is designed to prepare students for productive careers in biochemistry; its emphasis is training in research, and each student works closely with members of the faculty. In addition to the requirement for a Ph.D. dissertation based on original research, students are required to complete six advanced courses in biochemistry and related areas among the 135 total units required for the Ph.D. Selection of these courses is tailored to fit the background and interests of each student. A second requirement involves the submission of two research proposals which are presented by the student to a small committee of departmental faculty members who are also responsible for monitoring the progress of student curricular and research programs, and a journal club presentation. All Ph.D. students are expected to participate actively in the department's seminar program, and students are encouraged to attend and to present papers at regional and national meetings in cellular biochemistry and molecular biology. Teaching experience is an integral part of the Ph.D. curriculum and is required for the degree.

The Department of Biochemistry offers an M.S. degree only to students already enrolled in the Ph.D. program. Students should contact the Graduate Studies adviser for more details.

Those applying for graduate study should have at least a baccalaureate degree and should have completed work in cell and developmental biology, basic biochemistry and molecular biology, and genetics. Also required are: at least one year of university physics; differential and integral calculus; and organic, inorganic, and physical chemistry. The department is especially interested in those applicants who have research experience in biology or chemistry. Students must submit an application, including transcripts and letters of recommendation, by December (see web site) for admission in the following Autumn Quarter.

Applications should be submitted at http://gradadmissions.stanford.edu . Applicants are notified by March 31 of decisions on their applications. Stanford University requires scores from the Graduate Record Examination (GRE) (verbal, quantitative, and analytical), and applicants are encouraged to submit scores from the GRE Subject Test in biochemistry, biology, or chemistry. Applicants should take the October GRE exam.

All applicants are urged to compete for non-Stanford fellowships or scholarships, and U.S. citizens should complete an application for a National Science Foundation Predoctoral Traineeship. Students are provided with financial support to cover normal living expenses; Stanford tuition costs are paid. Applicants for admission to the department are considered without regard to race, color, creed, religion, sex, age, national origin, or marital status.

Postdoctoral research training is available to graduates who hold a Ph.D. or an M.D. degree. Qualified individuals may write to individual faculty members for further information.

At present, the primary research interests of the department are the structure and function of proteins and nucleic acids, the biochemistry and control of development processes, molecular motors and the cytoskeleton, the trafficking of proteins between membrane-bound organelles, the control and regulation of gene expression, bioinformatics/protein structure design, and the application of microarrays to problems in human health and disease.

© Stanford University - Office of the Registrar . Archive of the Stanford Bulletin 2011-12.   Terms of Use | Copyright Complaints

Stanford University PhD in Biochemistry

How much does a doctorate in biological chemistry from stanford cost, stanford graduate tuition and fees.

Does Stanford Offer an Online PhD in Biological Chemistry?

Stanford doctorate student diversity for biological chemistry, male-to-female ratio.

Of the students who received their doctor’s degree in biological chemistry in 2019-2020, 33.3% of them were women. This is less than the nationwide number of 47.5%.

Racial-Ethnic Diversity

Racial-ethnic minority graduates* made up 33.3% of the biological chemistry doctor’s degrees at Stanford in 2019-2020. This is higher than the nationwide number of 20%.

Majors Related to a PhD in Biological Chemistry From Stanford

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Peng Wu, MD, PhD

Wu, a clinical instructor of pediatric hematology and oncology, has received the 2024 Burroughs Wellcome Fund Career Award for Medical Scientists. The award supports physician-scientists at the early stages of their careers, enabling them to bridge advanced postdoctoral or fellowship training and faculty service. She will receive a five-year grant totaling $700,000 to study lineage programs that promote tumorigenesis in the developing liver.

About Stanford Medicine

Stanford Medicine is an integrated academic health system comprising the Stanford School of Medicine and adult and pediatric health care delivery systems. Together, they harness the full potential of biomedicine through collaborative research, education and clinical care for patients. For more information, please visit http://mednews.stanford.edu .

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Physiology Track Ph.D. Program

The Ph.D. track in Physiology prepares students for research and teaching careers in academic, industry, government, and other settings where broad-based biomedical sciences and physiology training are required.

The first year and a half of the Ph.D. program are largely devoted to course work. During the first year, students complete interdisciplinary course work in the Graduate Program in Biomedical Sciences https://graduate.ouhsc.edu/Graduate-Programs/PhD-Programs/Graduate-Program-in-Biomedical-Sciences , emphasizing molecular aspects of cell and organismal biology, along with research rotations.  Human Physiology is taken in the second year.

After successful completion of a qualifying examination, based on a proposed plan for their research, the Ph.D. candidate performs original research leading to a dissertation. During this time the student works closely with their mentor and receives guidance from an advisory committee. Completion of the program in four years is a realistic goal.

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